ON GRAMINEA 261 
Eriksson has divided this species into a number of forms which show 
certain differences, chiefly biological : they are given in what follows, but 
it must be understood that they are distinguished almost entirely by their 
host-plants. They all show the same scattered, brownish-orange uredo- 
sori. Some, it is true, have ewcidia, others are not known to have them, 
but this is a difference which time may remove; also Pole-Evans (1907) 
has shown that they present minute differences in the mode of germination 
of their uredospores. Many authors prefer to consider the biological races 
which follow as distinct species, but if that is done it is a mistake, which 
entails continual confusion, to retain the name P. dispersa for one of them. 
DiIsTRIBUTION: Europe, Asia Minor, North America, and in 
Australia (probably only introduced). 
(1) PUCCINIA SECALINA nov. nom. 
Puccinia dispersa (sens. strict.) Erikss. Ann. Sci. Nat. ser. 8, ix. 268, 
pl. xi, f. 1—6 (1899). Sydow, Monogr. i. 709. Fischer, Ured. 
Schweiz, p. 357, f. 261. Klebahn, Wirtswechs. Rostp. p. 237. 
‘ Adium Anchusae Erikss. et Henning, Getreideroste, p. 210. 
4. Asperifolia Pers. ; Cooke, Handb. p. 541; Micr. Fung. p. 197 p.p. 
Aicidiospores. Aicidia hypophyllous or often on the calyx 
and fruit, seated on rounded yel- 
lowish or reddish-yellow spots, 
cup-shaped, with an incised revo- 
lute margin; spores verruculose, 
orange, 20—26 p. 
fAKcidia on Anchusa (Lycopsis) 
arvensis, very rare, Shere, Folkes- 
tone, Eltham, etc, August; uredo- 
and teleutospores on Secale Cereale, / 
May—October. (Fig. 198.) 
The Brown Rust of Rye in the Fig. 198. P. secalina. Aicidia 
uredo- and teleuto-stages seems to be ate ace 
confined to that cereal. Eriksson and of Rye. 
Klebahn have proved that it can be : 
transferred from species of Anchusa to the Rye and vice-versdé, but not, 
they say, to other plauts. Plowright’s observation (Ured. p. 168) that 
the zcidium was produced on Anchusa by infection from a rusted bundle 
of wheat straw is discredited by them, but possibly without sufficient 
reason. The teleutospores are capable of germination as soon as they 
mature ; hence the ecidium is usually met with in August and September. 
