ZAGHOUANIA 331 
The description of the uredo- and teleutospores is after Fischer; the 
latter mature in autumn and germinate at once. According to him, the 
mycelium of the excidial stage is perennial in the rhizome, but Klebahn 
proved (Zeitschr. f. Pflanzenkr. 1907, p. 144) that the teleutospores infect 
the growing points of the rhizome in autumn, and produce the ecidia in 
the following spring. He could also infect other species of Pyrus (Aria, 
torminalts, scandica, Malus) from the Anemone; Fischer did the same for 
P. fennica and P. communis. In the Anemone nearly every leaf of the 
affected plant will be attacked, as well as the flower-shoots. The leaves 
become longer, narrower and of a paler green, and are borne on longer 
petioles. They are often divided into more segments than the normal 
leaves. Fischer remarks that, when the fungus appears on the sepals, the 
cells in the neighbourhood develop chloroplasts. 
The discovery of this hetercecism was due to Dietel and was confirmed 
by Klebahn ; previously the ecidium has been mistakenly attributed, by 
Soppitt, to Lndophyllum as £. leucospermum. The mode of germination 
of the spores will, of course, easily distinguish them: about this there 
seems to have been some misapprehension—Soppitt (Journ. Bot. 1893, 
p. 274) distinctly stated that the spores did not germinate with “ promy- 
celial” spores, but in the Trans. Brit. Myc. Soc. i. 84, 98, this is altered 
into the statement that the spores ‘germinate as do those of the (other) 
Endophylla.” 
Puccinia fusca lives upon the same plant (Anemone nemorosa) but 
affects its host in a different way (see p. 215). Meidium punctatum, 
a stage of Puccinia Pruni-spinosae, is found on A. nemorosa as well as on 
garden Anemones (A. coronaria), but differs in the character of the 
broader peridium and in having faintly coloured spores. Moreover, its 
spermogones are dark-coloured and are found on both sides of the leaf 
(see p. 207). 
ZAGHOUANIA Patouillard. 
Sorl erumpent, subpulverulent. .Aicidia (Peridermium) 
with a peridium which is irregularly lacerate above; margin 
slightly involute ; spores in short chains, soon seceding. Sper- 
mogones flask-shaped, with ostiolar filaments. Uredospores 
pedicellate, solitary. Teleutospores one-celled, ovoid, pedicel- 
late, with a slightly thickened hyaline and verruculose epispore, 
germinating as soon as mature; basidium four-celled, semi- 
internal; basidiospores nearly sessile. 
The description is founded upon that given by Dumée and 
Maire in Bull. Soc. Myce. Fr., 1902. The semi-internal basidium 
is characteristic. Dumée and Maire make a separate family, 
