370 THECOPSORA 
The connection of the spore-forms on these two hosts has been experi- 
mentally demonstrated by Klebahn, Tubeuf, and Fischer. The basidio- 
spores in spring (about the time of pollination of the Fir) infect the female 
flowers of the Spruce Fir, which are usually at the top of the high trees ; 
occasionally also the young shoots are affected, but they do not produce 
zcidia. The ecidia are developed in late summer, and mature on the 
fallen cones; their spores germinate in the following May, and then infect 
the leaves of the Bird Cherry, on which they produce uredospores in the 
summer and teleutospores in the autumn (Fischer, J.c. and Centralbl. f. 
Bakter. 2, xv. 1906, p. 227). The description of the teleutospores is 
taken from Klebahn and Fischer. As will be seen from the synonymy, 
the ecidium was originally classed among the Myxomycetes. The three 
stages appear in Cooke’s Handbook (according to the knowledge then 
prevailing) on three different pages, the ecidium from Appin, the uredo 
from some place in Scotland, and the “Melampsora” from Swanscombe, 
Kent (Cooke, 1865). It is also recorded on P. Padus in Yorkshire Fung. 
FI. p. 184, while the eecidium is recorded on ‘pine-cone scales” on p. 369. 
The uredo has also been found at Braemar, Aboyne, Perth, etc.; and the 
zecidium in Dumfriesshire. 
DISTRIBUTION: Europe. 
2. Thecopsora Galii De Toni. 
Caeoma Galii Link, Sp. Pl. ii. 21. 
Melampsora Galiz Wint. Pilze, p. 244. Plowr. Trans. Brit. Myc. Soc. 
i. 59. 
Puceiniastrum Gali Fischer, Ured. Schweiz, p. 471, f. 307. 
Thecopsora Galii De Toni in Sace, Syll. vii. 765. 
Uredospores. Sori scattered or gregarious, small, round, 
pulvinate, reddish, covered by the epidermis and by a peridium 
which opens at the summit with a pore; spores shortly ellipsoid 
or ovate, sparsely echinulate, orange-yellow, 17—20 x 14—16 yp; 
epispore colourless, without evident germ-pores. 
Teleutospores. Developed in the epidermal cells, forming 
httle dark-brown crusts, crowded, roundish, longitudinally 
septate into 2—4 cells, 21—24 x 21—32 yu; epispore rather 
thick, yellowish-brown, smooth, with an evident germ-pore at 
the upper and inner corner. 
On Galium verum (H. T. Soppitt), June—September, 1889. 
Very rare, 
It is reported, in continental Europe, as occurring on other species 
of Galium, also ou Sherardia and Asperula. 
