GREW 171 



side of the leaf. There is a tolerable figure of the 

 stomates of a lily-leaf (PI. XLVIII). It is singular 

 that Grew does not notice Malpighi's earlier discovery 

 of the stomates.^ 



Alternation of Floral Whorls 



The relative position of the parts of the flower is 

 explained ; they are not " filed one just over another, 

 but alternately," thus admitting of closer packing and 

 more complete protection of the inner organs (p. 164). 



The Flower 



In the flower Grew distinguishes the calyx (" empale- 

 ment"), corolla ("foliature"), and stamens ("attire"); 

 the pistil he does not consider a part of the flower, but 

 of the fruit ; this had been the view of Theophrastus and 

 also of the sixteenth-century botanists. He tells us 

 that the anthers ("semets") when they burst scatter 

 their powders, which are a "congeries, usually, of so 

 many perfect globes or globulets, sometimes of other 

 figures, but always regular" (p. 38). The uses of the 

 pollen Grew can only guess at ; it may serve " for food, 

 for ornament and distinction to us, and for food to other 

 animals " (pp. 39, 40). The " primary and private use " 

 of the pollen is discussed more fully in the fourth book ; 

 Grew is convinced that it must be great and necessary. 

 He has no experimental evidence to bring forward, but 

 makes one interesting remark. " In discourse hereof 

 with our learned Savilian Professor, Sir Thomas 

 Millington, he told me he conceived that the attire doth 

 serve as the male, for the generation of the seed. I 

 immediately replied that I was of the same opinion ; 

 and gave him some reasons for it, and answered some 



^ Supra, p. 155. 



