NARCISSUS. 



79 



to one parent and then to the other with chromic acid 

 and pyrogallic acid, the parental relationships being 

 reversed in these two reactions. 



(5) An early period of resistance followed by a com- 

 paratively rapid reaction is seen in the reactions with 

 chromic acid and pyrogallic acid — in all three starches in 

 the first and in the two starches in the second. 



(6) The earliest period at which the three curves are 

 best separated for differential purposes is in the sul- 

 phuric-acid reaction within the 5-minute period ; in that 

 with pyrogallic acid at 45 minutes; and in that with 

 chloral hydrate at 60 minutes. 



Eeaotion-intensities of the Hybrid. 



This section deals with the reaction-intensities of the 

 hybrid as regards sameness, intermediateness, excess, 

 and deficit in relation to the parents. (Table A 17 and 

 Charts D 399 to D 304.) 



The reactivities of the hybrid are the same as those 

 of the seed parent in the reactions with gentian violet and 

 chromic acid; the same as those of the pollen parent in 

 those with polarization, iodine, and safranin; the same 

 as those of both parents in none ; intermediate in none ; 

 highest in those with chloral hydrate, nitric acid, and 

 sulphuric acid, in all three being closer to the seed parent ; 

 and lowest in those with temperature and pyrogallic acid, 

 iu both being closer to the seed parent. 



The following is a summary of the reaction-intensi- 

 ties (10 reactions) : Same as seed parent, 2; same as 

 pollen parent, 3 ; same as both parents, ; intermediate, 

 ; highest, 3 ; lowest, 2. 



The seed parent, N. princess mwry, has from these 

 data exercised a far more potent influence than N. poeti- 

 cus poetarum on the properties of the starch of the 

 hybrid. 



Composite Curves of the Eeaction-intensitibs. 



This section treats of the composite curves of the reac- 

 tion-intensities, showing the differentiation of the 

 starches of Narcissus princess mary, N. poeticus poe- 

 tarum, and N. cresset. (Chart E 17.) 



The most conspicuous features of this chart are: 



(1) The very close correspondence in the curves, 

 both as to nearness and course. 



(2) In N. princess mary in comparison with the 

 other parent the higher reactions with gentian violet, 

 chromic acid, and nitric acid; the lower reactions with 

 polarization and iodine ; and the same or practically the 

 same reactions with chloral hydrate, pyrogallic acid, and 

 sulphuric acid. 



(3) In A^. princess mary the very high sulphuric- 

 acid reaction; the absence of any high reaction; the 

 moderate reactions with iodine, safranin, chromic acid, 

 and pyrogallic acid ; the low reactions with polarization, 

 gentian violet, temperature, and nitric acid ; and the very 

 low reaction with chloral hydrate. 



(4) In N. poeticus poetarum the very high reaction 

 with sulphuric acid ; the absence of any high reaction ; the 

 moderate reactions with polarization, iodine, safranin, 

 temperature, and pyrogallic acid ; the low reactions with 

 gentian violet, chromic acid, and nitric acid ; and the very 

 low reaction with chloral hydrate. 



(5) In the hybrid the very high reaction with sul- 

 phuric acid; the absence of any high reaction; the mod- 



erate reactions with polarization, iodine, safranin, and 

 chromic acid ; the low reactions with gentian violet, tem- 

 perature, pyrogallic acid, and nitric acid; and the very 

 low reaction with chloral hydrate. 



The following is a summary of the reaction-intensi- 

 ties (10 reactions) : 



18. Comparison's of the Starches of Narcissus 

 ABScissus, JST. poeticus poetarum, and ]Sr. will 



SCARLET. 



In histologic characteristics, polariscopic figures, 

 reactions with selenite, reactions with iodine, and quali- 

 tative reactions with the various chemical reagents the 

 starches of the parents and hybrid exhibit properties in 

 common in varying degrees of development, which collec- 

 tively in each case are distinctive, although all three 

 starches are very much alike. In histologic properties 

 the starches of the parents differ very little, and the 

 same is also true of the polariscopic figures and reactions 

 with selenite. In the iodine reactions no qualitative dif- 

 ferences were recorded. In the qualitative reactions with 

 chloral hydrate, chromic acid, pyrogallic acid, nitric acid, 

 and sulphuric acid there are properties in common and 

 also individualities. The starch of the hybrid ta com- 

 parison with the starches of the parents shows a closer 

 relationship to Narcissus ahscissus in the form of the 

 grains, the character of the hilum, the character of the 

 lamellffi, and the size of the larger grains ; but closer to 

 the other parent in the size of the smaller grains. The 

 eccentricity of the hilum is about the same in aU three 

 starches, and in the hybrid the lamellas are more distinct 

 than in the parents, and the hilum is not so deeply and 

 extensively fissured. In the polarization figures and 

 reactions with selenite the relationship is closer to A''. 

 atscissus. In the qualitative iodine reactions it is closer 

 to N. poeticus poetarum. In all of the qualitative reac- 

 tions with the chemical reagents peculiarities of both 

 parents are observed, but the resemblances are, on the 

 whole, closer to N. abscissus. Such differences as have 

 been recorded are only of a minor character. 



Reaction-intensities Expressed hy Light, Color, and Tempera- 

 ture Reactions. 

 Polarization : 



N. abscissus, low to high, value 43. 



N. poeticus poetar., low to high, somewhat less than in N. abscissus, 



value 40. 

 N. will scarlet, low to high, the same as in N. abscissus, value 43. 

 Iodine: 



N. abscissus, light to moderate, value 40. 



N. poeticus poetar., light to moderate, somewhat less than in N. 



abscissus, value 45. 

 N. will scarlet, light to moderate, the same as in N. poeticus poet- 

 arum, value 45. 

 Gentian violet : 



N. abscissus, light to moderate, value 33. 



N. poeticus poetar., light to moderate, somewhat more than in 



N. abscissus, value 35. 

 N. will scarlet, light to moderate, higher than in either parent, 

 value 37. 



