1914] DE VRIES— OENOTHERA LAMARCKIANA 355 



The plant which Potjrret called 0. grandiflora Lam. is repre- 

 sented on our pi. XVIII. It agrees wholly with the present 

 0. Lamarckiana Ser., and in all respects. It was fastened on its 

 sheet by the clerk of Pourret and consists of two flowering spikes 

 and two separate flowers. The stigma lobes are seen spread above 

 the anthers in the normal way. The specimens were picked at 

 the beginning of the flowering period and bear no fruits; obviously 

 they were main spikes. They will be recognized at once as 0. 

 Lamarckiana by anyone who has seen Hving cultures of this species. 

 As I have quoted in my Mutation theory (loc. cit.), Spach has 

 written on this sheet "Onagra vulgaris grandiflora Spach," which 

 remark also proves the identity with O. Lamarckiana Ser. The 

 printed label says "Collection de I'Abbe Pourret, extraite de 

 I'herbier legue par M. le Dr. Barbier en 1847." The main spike 

 measures about 40 cm., the smaller one about 20 cm. 



In my book I have also referred to a specimen of 0. suaveolens 

 Desf. At that time I did not know the Alabama species and 

 beheved that 0. suaveolens Desf. and 0. grandiflora Ait. were S3ai- 

 onyms, as almost all authors did. Therefore I used the two names 

 promiscuously. Last simuner, however, I cultivated, side by side, 

 0. suaveolens Desf. from Fontainebleau, collected by Dr. Blaring- 

 HEM, and O. grandiflora Ait. from Castleberry, Alabama, collected 

 by myself with Mr. Bartlett. They proved to be wholly different 

 species.^* So far as I know, the large-flowered Oenotheras, which 

 are now relatively common in the western departments of France, 

 all belong to O. suaveolens Desf., at least all the specimens and 

 cultures on which I based my opinion in igoi did. The specimen 

 of the Museum d'Histoire NatureUe, which I referred to especially, 

 has been described by Davis from a photograph which is repro- 

 duced on pi. 39 of his paper. Davis, who did not know the 0. 

 suaveolens as a separate species, called it the flotsam of the her- 

 barium (p. 529); it is, on the contrary, the authentic specimen of 

 Desfontaines, bearing on the label the name suaveolens written 

 by Despontaines himself. The smaller plant, fastened on the 

 same sheet, has another label, saying only O. grandiflora, and seems 

 to me to have been fastened on this sheet subsequently. The 



=* UOenothera grandiflora de I'herbier de Lamarck, loc. cit. 



