Davis: Lamarck's evening primrose 525 



istic of the younger flowering shoots of De Vries's Lamarckiana. 

 On the contrary the bracts are narrow and short-petioled and the 

 inflorescence is more open, in agreement with grandiflora. 



3. Buds. The buds are not stout as in the Lamarckiana of 

 De Vries and the sepal tips are much more attenuate, a distinctive 

 character of grandiflora. The form of the buds is exactly as in 

 grandiflora, and this character is one of the most important points 

 of agreement with this species. The pubescence on the sepals, as 

 described by M. Gagnepain, is short; there are not present the 

 numerous long hairs from papillae, which are characteristic of De 

 Vries's Lamarckiana. 



4. Flowers. The flowers have the very long and delicate 

 hypanthium characteristic of grandiflora. Miss Eastwood reports 

 that the petals in a bud dissected by her are entire. This is a 

 character typical of neither grandiflora nor the Lamarckiana of De 

 Vries, but as stated before, the petals of these forms vary so greatly 

 in the degree of their indentation that the character has little if any 

 taxonomic value. The length of the petals, between 3 and 4 cm., 

 is that of grandiflora and also of certain forms of De Vries's 

 Lamarckiana. The style extends beyond the tips of the anthers 

 so that the lobes of the stigma {s, in plate 37) are above the latter 

 and could not be pollinated in the bud. In these respects the 

 flower agrees with both grandiflora and the large-flowered types of 

 Lamarckiana in De Vries's cultures. 



5. Capsules. There are apparently no mature capsules on 

 the specimen, so direct comparisons are impossible. Since the 

 size and form of a capsule depends upon the development of the 

 ovules, i. e. upon whether or not the stigma has been fully polli- 

 nated, it is unsafe to accept statements of size unless there is 

 evidence that sufficient material has been examined. Lamarck's 

 statement that the capsules are short was probably based on 

 immature or partially pollinated capsules. His description of the 

 capsules as glabrous points to grandiflora; as stated before, the 

 capsules of De Vries's Lamarckiana are decidedly puberulent and 

 pilose. 



The characters of Oenothera grandiflora, which appear on the 

 herbarium sheet of Lamarck's plant, and those of the Lamarckiana 

 of De Vries's cultures may be more readily contrasted in the 

 following statement. 



