On^«; o/ CEnothera Lamarckiana De Vries. 237 



What we Jesire is further information on the composition of 

 the cultures of Carter and Company through the discovery of other 

 herbarium material of about the same date (1862) as that of the 

 sheet in the Gray Herbarium. It will be strange if Dr. Gray proves 

 to the only botanist who preserved specimens of the " Lamarckiana " 

 placed on the market by Carter and Company. There should be 

 made a persistent search through herbaria to bring forward any 

 specimen that may throw light upon the problem. This is a matter 

 in which it would seem that the British botanists can render a 

 great service. 



Another feature of tiie problem concerns the development of 

 some remarkable CEnothera floras in parts of England composed in 

 greater part of O. Lamarckiana or variants from this type. 



It is surprising how common have become Lamarckiana-Vike 

 forms in England. During the past three years several English 

 botanists have kindly replied to my request for seed of broad-leaved 

 forms of CEnothera biennis with green stems bearing red papillae at 

 the base of long hairs. From the seed sent to me I have grown 

 eleven different cultures in the hope of finding a type of biennis 

 which in the past I have greatly desired as a parent for a cross 

 with 0. grandiflora. All of these cultures have proved to be forms 

 essentially Lamarckiana as to habit, foliage and stem coloratipn, 

 but with smaller flowers than is usual for De Vries's plant. They 

 correspond very closely and some of them are indistinguishable 

 from the small-flowered races of Lamarckiana which I have 

 differentiated from material of De Vries.' They were not at all 

 the forms of biennis that I hoped to obtain, and could not be used 

 in my experimental work. 



The type of biennis that I wish should agree closely in 

 morphology with the biennis of the sand dunes of Holland, but it 

 should have the stem coloration characteristic of Lamarckiana, i.e., 

 the green portions of the stems should be punctate with red papillae 

 at the base of the long hairs. Tbe Dutch biennis has as far as we 

 know a clear green stem above, but it would not be surprising if a 

 form should be found with the stem coloration of Lamarckiana, 

 since races of American biennis occur differing only in the presence 

 or absence of red coloration in the stem papillae. Seeds of a type 

 agreeing with the Dutch biennis hav? been sent to me from the 

 botanical garden of Cambridge University and presumably the plant 



' See Davis, American Naturalist, vol. 46, p. 383, 1912. 



