150 BEADliEY MOOBE DAVIS 



Type VI. Differed from Type I in having plane leaves. 1 plant. 



Type VII. Differed from Type I in its narrow, plane leaves with red veins. 

 2 plants. 



Type VIII. Similar to Type VII except for intensely red sepals. 1 plant. 



Type IX. Leaves crinkled, stems clear green (papillae at the base of long hairs 

 not red as in Type I). 4 plants. 



Type X. Leaves crinkled, stem clear green as in Type IX, buds viscid-puberu- 

 lent only, not pilose. 38 plants. 



Type XI. Differing from Type X in that the lower leaves were red-veined. 

 1 plant. 



Type XII. Differed from Type X in having a red stem and the lower leaves 

 red-veined. 1 plant. 



To one familiar with hybrids of Oenothera the behavior of the 

 plant from Santa Cruz in this culture can have but one interpre- 

 tation. The plant could not have been representative of a pure 

 species. It must have had a germinal constitution of mixed 

 parentage (heterozygous), or, in other words, it must have been 

 of hybrid origin. No plant of an approximately uniform germinal 

 constitution (homozygous) would have thrown off such a variety 

 of types differing in such clear cut characters. 



This culture thus illustrates the necessity of testing experimen- 

 tally any plant which may be supposed to present evidence that 

 Oenothera Lamarckiana occurs in the American flora as a native 

 species. The discovery of the Santa Cruz plant suggested the 

 possibility that Lamarckiana grows wild in California. The cul- 

 ture from its seeds showed that' the plant was hybrid in character 

 and consequently was not representative of a native species. 

 Its history is not known but we suspect that the plant was a 

 garden escape from Lamarckiana (frequently grown for ornamen- 

 tal purposes) which had hybridized with some other species of 

 the neighborhood. It was certainly not pure Lamarckiana be- 

 cause of the very great number and variety of the different types 

 thrown off in the culture. 



A brief description of Oenothera Lamarckiana De Vries together 

 with some photographs illustrating its most important characters 

 will be given for those who are not familiar with the plant. 



1. Rosettes. The mature rosette (fig. 2), 4 to 5 dm. broad, consists of broadly 

 elliptical or spatulate leaves with sinuate margins, irregularly toothed below, and 

 very strongly crinkled. There are forms of Lamarckiana with broader leaves. 



