[Vol. 1 

 390 ANNALS OF THE MISSOURI BOTANICAL GARDEN 



in pi. 21 fig. 11. It will be seen that there is no indication of a 

 rosette, and the branching is quite different from that of CE. 

 mut. rubrinervis. In many cases, however, a rosette is formed. 

 "When the rosette is omitted the branching is changed. Plate 

 21 fig. 10 shows on a larger scale another individual in flower. 

 The stem-leaves differ from those of (E. mut. rubrinervis in being 

 narrower, more pointed and smoother. 



In this race the red papillae on the stem were very numerous, 

 and the buds likewise were shghtly more red than in CE. mut. 

 rubrinervis. The modal color pattern of the whole population 

 was 5 as in (E. mut. rubrinervis, but plants with their mode at 

 7 were much more numerous than in the latter (see Gates, '11, 

 p. 351). The race as a whole inherited the capacity for pro- 

 ducing a slightly greater amount of pigment. The ovary usu- 

 ally bore many long hairs arising from red papillae; on the hy- 

 panthium were few long hairs from sUght green mounds; and 

 on the bud cone scattered long hairs from conspicuous red 

 papillae. In occasional buds, when the color pattern was only 

 S, the green papillae were more numerous. In addition to the 

 color pattern of the sepals there was usually weak red on the 

 hypanthium.i 



The same conditions as regards pigmentation have been main- 

 tained in later generations. The plants were, however, by no 

 means uniform in all respects, and this was not to be expected 

 since they were derived from one individual of a freely inter- 

 crossing population. Plate 21 fig. 9 represents a rosette of 

 one of the F2 plants. The latter differs obviously from the one 

 represented in pi. 21 fig. 8, but the race retained in this and 

 subsequent generations the long, narrow, smoothish leaves as 

 well as the pigmentation. The various Fs and F4 families, each 

 derived from a selfed individual, produced sub-races differing 

 more of less from each other and varying within narrower limits. 

 It does not appear that the MendeHan theory of the sorting out 

 of factors, or "genes," affords an adequate explanation of all 

 these phenomena. 



' Since this condition of bud-pigmentation resembles that obtained in certain Fj 

 and Fj hybrids of (E. mut. ruhricalyx and (E. grandifiora (see Gates '14), it is possi- 

 ble that it may have arisen in a similar way, i. e., by the appearance of a red-budded 

 mutation which subsequently crossed with other species, in which crosses some blend- 

 i ng of pigmentation occurred giving rise to the present condition. 



