1914] 



GATES — SOME (ENOTHEEAS FROM CHESHIRE AND LANCASHIRE 393 



two on the right in pi. 22 fig. 18 were greatly overgrown and 

 were far larger than ever appear even in (E. mut. gigas. These 

 forms have not been sufiiciently studied since to give an ade- 

 quate account of them. 



It will be obvious that the forms described here under the 

 names multiflora, multiflora elliptica, ruhrinervoides, tardiflora 

 and rubritincta are not pure species or even true-breeding races. 

 They are undoubtedly as diverse from each other as average 

 species, however, and many systematic species if bred experi- 

 mentally would probably not breed true within narrower limits 

 than these races have done. One feature of interest attaching 

 to these races is the fact that the main type persists essentially 

 unchanged, though various mutants and heterozygous forms 

 are thrown off. The behavior is not, in the main, like the Men- 

 delian process of recombination. Repeated selfing of each race 

 usually decreases its variability by eliminating various hybrid 

 elements. But this process does not extend to the basal differ- 

 ences between the races, which, as we have seen, remain as 

 unlike as they were before. In this aspect the hereditary 

 behavior of these races resembles that of (E. Lamarckiana. 

 But there are a number of differences which I need not fully 

 consider. Thus (E. multiflora gives rise to its variety elliptica 

 much as though it were split off from a heterozygous condi- 

 tion, and the variability of rubritincta in leaf -width, as well as 

 its production of numerous dwarfs, is unlike anything in the 

 behavior of (E. Lamarckiana. 



Many other equally distinct types were derived from this 

 locality (see, e. g., pi. 22 figs. 18, 19), but they have not been 

 cultivated in subsequent generations. 



CE. LAMARCKIANA FROM ST. ANNE'S 



In 1910 I obtained seeds from a colony of (E. Lamarckiana 

 growing by the Manchester Children's Hospital Convalescent 

 Home, at St. Anne's-on-Sea. Many of these were found in 

 later cultures to agree exactly with the Lamarckiana of de Vries 

 except in the red color pattern of the sepals. I was formerly in- 

 clined to lay little stress on this difference but there is no doubt 

 that it is inherited. The fact therefore remains that a precise 

 duplicate for de Vries's race of (E. Lamarckiana is relatively 



