191«] 



GATES — SOME CBNOTHERAS FROM CHESHIRE AND LANCASHIRE 395 



squarish). Two other plants agreed exactly with (E. Lamarck- 

 iana except in the buds. The petals were 35 mm. long x 48 

 mm. broad, emarginate, anthers reaching nearly to top of stigma 

 lobes, sepals green and with the same pubescence as in (E. 

 Lamarckiana, from which these two plants therefore differed 

 only in the somewhat smaller flowers and shorter style. One 

 mut. nanella also occurred in this culture, and several other 

 sUghtly aberrant individuals, including a plant with broadly 

 elliptical foUage. The "Lamarckiana foliage" was also more 

 variable than in cultures from de Vries, this no doubt being due 

 to the continued inbreeding in the latter case. 



It will be understood that the new forms described here are 

 scarcely to be looked upon as "new species" according to the 

 usual interpretation at the present time. They merely represent 

 a partial analysis of a complex interbreeding colony of forms, 

 and their variability is one of their most interesting features. 

 Nearly all if not all the differences observed are inherited, how- 

 ever, and the mutations can in many instances be separated 

 from the characters arising through hybridization. The forma 

 are, moreover, as distinct from each other as many species of 

 (Enothera. 



In conclusion, I am indebted to the Missouri Botanical Garden 

 and the John Innes Horticultural Institution for the facilities 

 provided for growing the plants, and to Mr. E. J. Allard for 

 several of the photographs. A portion of the expenses of my 

 second visit to Lancashire was defrayed by a grant from the 

 Royal Society. 



LiTEEATURE CiTED 



Bailey, C. ('07). De Lamarck's evening primrose {(Enothera Lamarckiana) on the 



sandhills of St. Anne's-on-the-Sea, North Lancashire. (Address, Annual Meeting 



Manchester Field Club.) 1-28. pi. 1-6. 1907. 

 Gates, R. R. ('10). Abnormalities in (Enothera. Rept. Mo. Bot. Gard. 21: 175-84. 



Vl. B9-31. 1910. 

 , ('11). Studies on the variabihty and heritabiUty of pigmentation in CEno- 



thera. Zeitschr. f. induct. Abst.- u. Vererbungsl. 4: 337-72. pi. 6. f. 1-5. 



1911. 

 , ('12). An onagraceous stem without internodes. New Phytologist 11: 



50-53. pi. Z-3. 1912. 

 , ('13). A contribution to a knowledge of the mutating cenotheras. Trana. 



Linn. See. Bot. 8: 1-67. pi. 1-6. 1913. 



