742 LXIT. EUBIACE^. 



Order LXIV. RUBIACE^. 



Calyx-tube adnate to the ovary ; the limb entire or with as many teeth lobes 

 or divisions as lobes of the corolla, rarely more, fewer or none. Corolla gamo- 

 petalous, inserted round the epigynoiis disk ; lobes 4, 5 or sometimes more, rarely 

 only 8, either imbricate (often contorted) or valvate in the bud. Stamens as 

 many as lobes of the corolla, alternating with them and inserted in the tube ; 

 anthers versatile, with parallel cells opening longitudinally. Ovary inferior, 2 or 

 more-celled, with 1 or more ovules in each cell, rarely 1-celled with parietal , 

 placentas, or reduced to one 1 -ovulate cell ; style more or less divided into as 

 many stigmatic lobes as carpels to the gynoecium, or undivided with a thickened 

 entire or notched stigma. Fruit a capsule, drupe, berry or indehiscent nut. 

 Seeds with a fleshy or horny albumen, and rather small straight embryo with 

 flat cotyledons, or rarely with little or no albumen and cylindrical embryo with 

 semiterete cotyledons. — Trees, shrubs, herbs, or rarely climbers. Leaves opposite 

 or whorled. Stipules interpetiolar, either free or connate with the petioles in a 

 sheath bordered by cilia or leaf-like lobes, or with one or two points on each side, 

 or connate witfiin the petioles in a short sheath or ring round the stem. 

 Inflorescence various, usually more or less cymose, axillary or terminal. Flowers 

 occasionally polygamous or unisexual. Species chiefly tropical and subtropical. 



Series A. — Ovules numerous in each cell (rarely in a few HedyotuJem). 



Tbibe I. Na.UCleeSB. — Flowers collected into dense globose heads. Corolla funnel-shaped : 

 stigma simple. 



"Ovaries conAne.nt; fruits fanning a globose fleshy mass ; corolla-lobes imbricate in Imd. 



Ovary 2-ceUecl, ovules numerous 1. SvncocF.nnnjs. 



""Ovaries free or nearly so; fruits free, capsular. 

 Flowers ebracteolate. 



Corolla-lobes imbricate 2. Nauclea. 



Corolla-lobes valvate, climbers with hooked peduncles 'i. XJncabia. 



Tribe II. Zlondeletlete. — Calyx-lobes twisted in bud. Fruit capsular, 2-celled. Seeds 

 angular. Shrubs or trees. 

 Flowers simply paniculate . ... . .4. Wendlandia. 



Tribe III. KedyotldeaB. — Corolla-lobes valvate in bud. Fruit a loculicidal or septicidal 



2 rarely i-celled capsule, or of 2 or i dehiscent or indehincent cocci, rarely altogether indehiscent. 



Seeds small or minute, rarely winged. Herbs, rarely shrubs. 



"Fi^uit '2-celled, usually indehiscent, stipules entire. 



Corolla funnel-shaped, lobes 4 or 5-toothed 5. Destblla. 



""Fruit oblong, subglobose, or orbicular, 2 rarely i-celled, usually dehiscent; cells many rarely 



1-seeded. 



Capsule loculicidal or septicidal, rarely indehiscent, with contiguous calyx- 

 teeth, seeds usually angular 6. Hedyotis. 



Capsule loculicidal above the remote calyx-teeth, rarely indehiscent ; seeds 

 minute, angular 7. Oldenlandii. 



Stamens 4. Capsule f-superior. Corolla rotate, mareesoent, retaining the 



stamens . . 8. Stn'aptan'tha. 



"""Fniit broadly didymously obcordate, of 2 compressed spreading lobes, loculicidal above the 



calyx. 



Flowers crowded on the upper side of the branches of a pedunculate cyme . 9. Ophior^hiza. 



Tbibe IV. musssendese. — Corolla-lobes valvate in bud. Fruit fleshy or drupaceous, inde- 

 hiscent. Seeds minute. 

 Corolla-lobes 4, induplieate-valvate. Anthers cohering in a tube. Fruit a 



berry, seeds numerous 10. Abbottia. 



Tkibe V. Gardeniese. — Corolla-lobes twisted in bud. Fruit berried, fleshy, or dry. Seeds 

 usually large; cotyledons often foliaceous. 



"Seeds usually many in each cell. 

 Inflorescence terminal, flowers 2-sexual, stigma fusiform . 11. Webeea. 



Inflorescence usually axillary, stigma fusiform. 



Ovary 2-celled 12. Eandia. 



Ovary 1-celled 13. G.akdenia. 



""Style-arms 2, free. Ovary 2-celled, Seeds few in each cell. Inflorescence 'axillary. 

 Calyx calyculate, limb truncate or 2 to 5-lobed, flowers fascicled, rarely 



cymose . , , ... U. Diplospora. 



