194 HEREDITY AND EVOLUTION IN PLANTS 



our museums," says Darwin, "is absolutely as nothing 

 compared with the number of generations which must 

 have passed away during a single formation." The 

 meagerness of the record is, of course, due in part to the 

 relatively small area explored in proportion to the whole ; 

 but there are other reasons much more serious, because 

 they represent opportunities lost forever. Among them 

 are metamorphosis, explained above, and the fact that 

 many of the organisms of the past were composed wholly 

 or largely of soft tissues, which were entirely destroyed, by 

 decay or otherwise, in the process of rock-formation. 

 Such plants, for example, as Spirogyra and many other 

 algae, the fleshy fungi, and, among animals, jelly-fish, 

 earthworms, and others, would form fossils only under 

 exceptionally favorable circumstances, if at all. 



But there is an even more effective cause of oblitera- 

 tion of the fossil record in the long-continued erosion and 

 denudation represented by unconformity in the rock 

 strata. In many cases only a small proportion now re- 

 mains of the thickness of a rock stratum originally de- 

 posited, and all traces of the plant and animal life that 

 may have existed on the denuded area have thus been ob- 

 literated forever. These blank intervals between suc- 

 cessive periods were of vast duration. 



"I look at the geological record," said Darwin, "as a 

 history of the world imperfectly kept, and written in a 

 changing dialect; of this history we possess the last 

 volume alone, relating only to two or three countries. 

 Of this volume, only here and there a short chapter has 

 been preserved; and of each page, only here and there a 

 few lines. Each word of the slowly changing language, 

 more or less different in the successive chapters, may 



