THE MORPHOLOGY AND SYSTEMATIC POSITION OF 

 CALYCULARIA RADICULOSA (Steph.) 



By Douglas Houghton Campbell^ Professor of Botany 



THE classification of the so-called anacrogynous Jungermanniales, an 

 important group of liverworts, is at present in a very unsatisfactory 

 condition, and much remains to be done before the true relationships 

 of the members of this group can be satisfactorily settled. A recent attempt 

 has been made toward a better classification of the liverworts by Cavers^ 

 and this is a distinct advance upon the classifications which have heretofore 

 been accepted. There are, however, a number of forms whose relationships 

 are still by no means clear, and among these is a rare liverwort from Java 

 originally described as Calycularia radiculosa. Schillner^ speaks of the 

 plant as an extremely rare one, but during a stay in Tjibodas where the 

 plant had been collected before, the writer succeeded in finding it a number 

 of times. The plants did not grow in large masses but were associated with 

 various other liverworts growing on the trunks of trees. The general aspect 

 of the plant (see Figs. 1 and 2) is very much like that of the creeping 

 forms of the genus Blyttia. Pallamcinia (Blyttia) Levieri, a common species 



Fig. 1. Three male plants of Calycularia 

 radiculosa. Steph. x 3. $ , antheridial recep- 

 tacle. 



iThe inter-relationships of the Bryophyta. New Fhytologist Rejprint, No. 4, 



1911. 



2 Die Hepaticae der Flora Von Buitenzorg, 1900. 



