XIII 



LYCOPODINEM 



489 



atum, although the essential structure is much the same. It is 

 a fusiform structure, with a terminal mass of short, irregular 

 lobes covered with the reproductive organs. Between the ter- 

 minal generative portion and the sterile fusiform body of the 

 prothallium, there is a meristematic zone, corresponding to that 

 in L. clavatum. The oldest reproductive organs are at the 

 centre of the generative area, the youngest are next the zone of 

 meristematic tissue. 



L. Selago closely resembles L. phlegmaria in the structure 

 of the gametophyte, and there are similar paraphyses formed 

 among the reproductive organs. 



L. inundatum, as was pre- 

 viously shown by Goebel, .be- 

 longs to the type of L. cer- 

 nuum, and Phylloglossum 

 (Thomas (i)) seems to be 

 very much like L. cernuum, 

 in the structure of the game- 

 tophyte. 



The gametophytes of all 

 species are normally dioe- 

 cious, but the antheridia 

 usually develop first. 



The Sexual Organs 



Fig. 284. — A, Lycopodium clavatum, gameto- 

 phyte, X3; B, L. annotinum, old game- 

 tophyte, with young sporophytes, sp, at- 

 tached, X3: C, gametophyte of L. com' 

 planatum, X3 (after Bruchmann). 



The sexual organs of all 

 investigated species of Lyco- 

 podium are very similar, and 

 resemble those of the eusporangiate Ferns and Equisetum. 

 As in these forms the antheridium mother cell divides first by a 

 periclinal wall into an outer and inner cell, the latter giving 

 rise immediately to the sperm cells. In the outer cell the divi- 

 sions are much like those in Marattia, but the opercular cell 

 does not become detached as in these, but is broken through 

 as in the Polypodiacese. In L. phlegmaria the outer wall is 

 often in places double, as not unfrequently is the case in the 

 Ophioglossaceas. The spermatozoids are almost straight ob- 

 long bodies with two cilia, like those of the Bryophytes (Fig. 

 283, G). The vesicle, which usually remains attached to the 

 spermatozoids of most Archegoniates, here is almost always 



