APPENDIX 621 



The species of Megaceros are mostly tropical, and they are especially 

 common in certain parts of the Malay Archipelago. The writer has 

 collected them at various stations in Java, Sumatra, Borneo, and 

 Luzon. Some of the species are very large and conspicuous, and occur 

 in masses covering the rocks in stream-beds and similar localities. 

 Others grow on rotten logs, and less commonly on the ground. 



The thallus usually closely resembles that of the larger species of 

 Anthoceros, and the apical growth in the species investigated by the 

 writer is exactly the same. The most obvious difiference is the 

 presence of several chromatophores in the cells, sometimes as many 

 as twelve having been observed in the inner cells. Usually no 

 pyrenoid can be recognized, and the chromatophores are much like 

 those of the higher plants. 



The antheridia are large, and borne singly as in Dendroceros or 

 Anthoceros Pearsoni. 



The sporophyte in its earlier stages is most like that of Dendroceros, 

 but there is a much larger development of the sporogenous tissue, 

 which suggests the condition found in Notothylas. The spores at 

 maturity contain chlorophyll, a condition found also in Dendroceros, 

 but not in Anthoceros, and the elaters have spiral thickenings as in 

 Dendroceros. Like the latter, stomata are absent. 



Megaceros is thus a sort of S5mthetic type, combining characters 

 found in all three of the other genera. (See Campbell (30).) 



P. 148. The writer has investigated two species of Dendroceros 

 from Java (Campbell (30, II) ), which agree closely with the other 

 species that have been examined. 



P. 156. Lang (7) states that in a species of Notothylas from Singa- 

 pore (probably N. Breutelii), while the early stages of the embryo 

 agree with the other Anthocerotacese, and the primary sporogenous 

 tissue originates from the amphithecium, the upper portion of the 

 columella develops spores, so that the latter arise in part from the 

 endothecium. A similar condition, but less marked, was found by the 

 writer in N. Javanicus. (Campbell (30, II).) 



P. 159. While there are certain similarities between the young 

 sporophyte of the Anthocerotaceae and such Liverworts as Sphmro- 

 carpus, Cyathodium and especially Fossomhronia, the fact that the 

 primary sporogenous tissue in the Anthocerotales always arises from 

 the amphithecium, while in all other Liverworts it is developed from 

 the endothecium, would seem to be a radical difference. Cavers, 

 however, thinks that the differences between the Anthocerotaceae and 

 the other Liverworts are not suflScient to warrant removing the Antho- 



