APPENDIX 633 



P. 298. All of the forms of Angiopteris have been referred by some 

 writers to a single species, A. evecta, but there is no question that 

 there are a number of well-marked species, although probably some 

 of the species recognised by De Vriese (i), should be eliminated. 



P. 300. The genus Macroglossum was first described by Copeland 

 (i), from material sent from Sarawak in Western Borneo, where it 

 has been collected at several points. A form of this, probably a 

 second species, has been cultivated in the Botanical Garden at Buiten- 

 zog, Java, xmder the name Angiopteris Smithii. Macroglossum has 

 also recently been reported from Sumatra. Macroglossum, unlike 

 Angiopteris, has simply pinnate leaves, and the structure of the sporan- 

 gium is more like that of Archangiopteris, to which it is more nearly 

 related than it is to Angiopteris. (See Campbell (35, 36).) The 

 type, M. AlidcB, is a large fern with leaves sometimes nearly four 

 metres in length. 



P. 300. Some species of Danaa, e.g., D. elliptica, have an upright 

 rhizome, and the leaves arranged spirally. 



P. 300. Chlorophyll may develop under certain conditions in 

 the gametophyte of Ophioglossum (see Bruchmann (5), Mettenius 

 (2)). 



P. 301. The young embryo of Ophioglossum Moluccanum, re- 

 sembles very closely that of Anthoceros. 



P. 303. The recent studies of the writer on the embryology of the 

 Marattiaceae and Ophioglossacese show a much greater similarity 

 between them than was supposed to be the case. (See Campbell 



(33)-) 



P. 304. The reasons for the assumption of a direct relationship 

 between the Ophioglossaceae and Marattiaceae have been given at 

 length elsewhere. (See Campbell (^s).) The conclusions reached 

 may be briefly summarised. " From some form allied to the existing 

 species of Ophioglossum the whole Fern-series is descended. In this 

 series the leaf is the predominant organ, the stem, at first, being of 

 quite subordinate importance. This ancestral Fern was monophyUous 

 and the original leaf was a sporophyll, perhaps without any definite 

 sterile segment. 



From this central type it may be assvuned that several divergent 

 lines of development arose, of which only isolated fragments have 

 persisted to the present time. 



The Marattiaceae, as they now exist, probably do not represent a 

 single unbroken line of descent, but show evidences of a multiple 

 derivation from the primitive stock. The point of contact with the 



