PHYLLODOCE. 75 



the ciliated genital funnel as open in front and closed behind, and intimately con- 

 nected with the segmental organ, which is closed internally, branched, and furnished 

 with solenocytes. 1 At maturity an opening is found at the point of fusion of the two 

 structures, so that the ripe products escape ventrally. 



Nervous system. — Cephalic ganglia with antero-superior and postero-inferior lobes, 

 the latter, the antennary centre, most bulky. The antero-superior centres give origin 

 to the stomato-gastric which receive an increment from the oesophageal connectives. 

 Ventral chain with fused ganglia, the connecting trunks being separate, and the first pair 

 larger than the others, and apparently formed by the fusion of three. 



Reproductive organs developed from ccelomic epithelium, and shed into the perivisceral 

 space. Goodrich and Jourdan think that the ova escape by the segmental pores. 



Gravier 2 describes foot-glancls debouching near the segmental aperture ventrally. 



Ehlers (1 868) characterises Phyllodoce, Savigny, thus : " Body elongated, generally 

 flattened, distinctly segmented ; head with four tentacles, the first two segments with four 

 pairs of tentacular cirri, and often with a rudimentary foot ; the remaining segments 

 distinct, with simple feet bearing a fan-like group of jointed bristles and leaf -like dorsal 

 and ventral cirri." 



Grube (1879) made two groups of Phyllodoce, viz. : (1) those with the head more or 

 less heart-shaped, having a deep dimple posteriorly (P. laminosa and P. groenlandica) ; 

 and (2) those with the head heart-shaped or rounded, but with the posterior border only 

 a little notched (P. maculata and P. citrina). 



De St. Joseph (1888), after a critical examination of the classification of the 

 " Phyllodoces," adopts the following genera : (1) Genetyllis, Malmgren. Achetous buccal 

 segment with four pairs of tentacular cirri ; (2) Phyllodoce sans str., Czerniavsky. 

 Achetous buccal segment with two pairs of tentacular cirri, other two pairs in the following 

 setigerous segment; (3) Anaitis, Malmgren, nee Claparede. Buccal segment achetous, with 

 three pairs of tentacular cirri, second segment with one pair, and with bristles; (4) 

 Garobia, De Quatrefages, Marenzeller, rev. {Anaitis, Clap., Phyllodoce, Mgrm., Anaitides, 

 Czerniavsky). Achetous buccal segment with one pair of tentacular cirri, fused or not 

 with the following segment, which has two pairs of tentacular cirri and bristles ; or the 

 achetous buccal segment with two pairs of tentacular cirri, fused or not with the following 

 segment, which has one pair of tentacular cirri and bristles ; third segment with one pair 

 of tentacular cirri, a setigerous lobe more or less developed, and a foliaceous ventral cirrus. 



Cunningham 3 says: "In Phyllodoce no well-marked canals can be distinguished. 

 The (nerve) cords are widely separated from the epidermis." 



Leschke 4 procured trochophores of Phyllodocidse in the Bay of Kiel in July, and 

 older forms in September, October, and November. 



1 c Q. Journ. Micros. Sci/ vol. 43, n.s., p. 706. 



2 An interesting account of the general structure of the group is given by the author in his 

 thesis, "Kecherches sur les Phyllodociens," Lille, 1896. 



3 Op. cit., p. 270, 1888. 



4 ' Wiss. Meer. Biol. Anstalt, Helgol./ n.f., 7 Bd., p. 117, 1903. 



