644 ^ nderson and Schafer. — Species Hybrids in Aquilegia. 



and the results are presented in Figs. 3 and 4. Two characters were chosen 

 for study, spur length and lamina length. In order to remove the effects 

 of hybrid vigour, in so far as they cause changes in size, comparative ratios 



2.1 1 



% 



\l.5 



•I- /.3 



■%. -3 

 ^ .7 



K 



\ 



k 



.5 



Cross /^f/ 



K 



2 3^5678 



Fig. 3. Analysis of the ratio lamina length/lamina width in first generation hybrids. The 

 vertical lines represent the difference between the parents in respect of this character. The arrow 

 indicates the position of the Ki hybrid on the same scale, i. A vulgaris compactay. A. fonnosa. 

 i. A caeruleax A. vulgaris. 3. A. ecalcaratax. A. chrysantha. 4. A. ecalcarataxA. vulgaris 

 compada. 5. A. chrysantha x A. vulgaris ' China lilue ' 6 A. vulgaris x A. Jaeschkani. 

 7. A. vulgaris x A. chrysantha. 8. A. vulgaris -n A. canadensis. 



were used. Since the width of the lamina at the orifice is a relatively con- 

 stant character it was used as a basis for comparison, the actual ratios 

 studied being: 



1. Length of lamina/width of lamina. 



2. Length of spur/width of lamina. 



Figs. 3 and 4 represent for each of the eight crosses the position of the 

 parent species on the two scales, the theoretical intermediate positions for 

 the first generation hybrid, and its actual position (at the right). 



The tables present in a more exact way what has already been stated 

 in general terms. In every case where the hybrid is not at the theoretical 

 midpoint between the parent species, the shift is in the direction of medium 

 length spurs and wide laminae. In other words, in species crosses in Aqui- 

 legia there is a general dominance of medium length spurs over very long 

 or very short spurs, and of wide laminae over long laminae. This is without 

 any evident relation to the particular species used, and is irrespective of the 

 direction in which the crosses are made. 



The regularity of the phenomenon seems to be adequately established 

 even by these preliminary studies. For its accurate interpretation further 

 work with more authentic material will be nece.ssary. Were it not for the 

 results obtained with A. ecalcarata one might ascribe the phenomenon to 



