Chap. II. HYBEID VERBASCUMS. Y3 



capsule. So that out of the eight possible unions be- 

 tween the two forms of these two species, six were utterly- 

 barren, and two fairly fertile. We have seen also 

 the same sort of extraordinary irregularity in the re- 

 sults of my twenty different crosses (Tables 14 to 18), 

 between the two forms of the oxlip, primrose, and cow- 

 slip. Mr. Scott remarks, with respect to the results 

 of his trials, that they are very surprising, as they 

 show us that "the sexual forms of a species manifest 

 in their respective powers for conjunction with those 

 of another species, physiological peculiarities which 

 might well entitle them, by the criterion of fertility, 

 to specific distinction." 



Finally, although P. veris and vulgaris, when crossed 

 legitimately, and especially when their hybrid offspring 

 are crossed in this manner with both parent-species, 

 were decidedly more fertile than when crossed in an 

 illegitimate manner, and although the legitimate cross 

 effected by Mr. Scott between P. auricula and hirsuta 

 was more fertile, in the ratio of 56 to 43, than the 

 illegitimate cross, nevertheless it is very doubtful, 

 from the extreme irregularity of the results in the 

 various other hybrid crosses made by Mr. Scott, whether 

 it can be predicted that two heterostyled species are 

 generally more fertile if crossed legitimately (i. e. when 

 opposite forms are united) than when crossed illegiti- 

 mately. 



Supplementary Note on some wild hybrid Verhascums. 



In an early part of this chapter I remarked that few 

 other instances could be given of a hybrid spontane- 

 ously arising in such large numbers, and over -so wide an 

 extent of country, as that of the common oxlip ; but per- 

 haps the number of well-ascertained cases of naturally 



