ATTACHMEXT OF THE AXTHER 



63 



and the production of a secondary or "false"' partition separating each 

 theca longitudinally into two locelli. This implies a four-locellate 

 condition of all anthers (Fig. 13S). Ordinarily this condition is not 

 permanent, the false septa more or less completely disappearing after 

 the formation of the pollen, lea\"ing the mature anther two-celled, or 

 this condition is brought about in other ways. 



Attachment of the Anther. — The Adnaie Form. — It is furthermore 

 assumed that the filament is normally continued along the back of 

 the anther in the relation of the midrib of the formative leaf. This 

 form of attachment is called Adnate (.Fig. 126). 



Fig. 126. Adnate anther of Maffnolia. 127. An incumbent anther. 12S. Twisted anther of Ceiba. 

 129. Versatile anther of Oenothera. 130. Innate anther of Saiiffuinaria. 131. Renifortn confluent 

 anther of Malr<i. 132. Horizontal confluent anther of Penislemon, 133. Sagittate anther of Taber- 

 naimontana. 



Incumbent Form. — It may be attached only at some point upon the 

 back (DorsLfixed). Of this there are two forms. In one (Fig. 127) the 

 anther is rigidly fixed, its lower portion close to and parallel ^"ith but 

 free from the upper portion of the filament, the Incumbent form. 



]'ersatUe Form. — In the other, it moves freely upon the pivotal 

 point of attachment (Fig. 129\ the Versatile form. Rarely the anther 

 is ^\Tapped or twisted about its filament (Fig. 12S^. 



Innate Form. — The continuation of the filament, instead of being 

 along the back, may be centrally up through the base and between the 

 thecae (Fig. 130\ the Innate form. 



Sagittate Form. — The lower portions of the thecae may be separated 

 from one another and from the connective i,Fig. 133\ the Sagittate form. 



Reniform and Horizontal Forms. — The sagittate condition is some- 

 times extreme, the anthers becoming more or less reniform (.Fig. 131) 

 or semicircular, or they may even become horizontal (Fig. 132\ This 



