CCELENTERATA : HYDROZOa. 1 1 1 



forms — doubtfullybelonghig to the sub-class — there is reason 

 to believe that the hydrosoma was fixed. The polypites are 

 never separated from the coenosarc by any partition. In many 

 cases the hydrosoma was strengthened by a solid chitinous rod, 

 the " solid axis," somewhat analogous to the chitinous rod re- 

 cently described by Professor AUman in the singular Polyzoon, 

 Rtiabdopleura. 



From the above definition, it will be seen that the nearest 

 living allies to the Graptolites are the Sertularians. In point 

 of fact, if we do not insist upon the presence of a " solid axis " 

 as part of the definition, the Graptolites differ from the Ser- 

 tularians in no essential point, save that the hydrosoma is 

 alwaj^'S attached in the latter, and was certainly free in the 

 most typical examples of the former. Indeed, certain forms 

 at present placed among the Graptolites — such as Ptilograpsus 

 and Dendrograpsus — are so similar to some living Sertularians, 

 that it might be well to remove them altogether from the 

 Graptolitidce, and to regard them as extinct representatives of 

 'Oa& Sertularida. 



As" regards the value of the "solid axis" as an elenient in 

 defining Graptolites, we fear that much stress cannot be laid 

 upon its presence or absence. It is true that it is present 

 in all the most characteristic members of the sub-class, but 

 it seems to be certainly absent in some — e.g., in HetioUtes 

 Geinitzianus, and in all species of Rastrites — and there do 

 not seem to be sufficient grounds for excluding these from 

 the Graptolitida on this account alone. 



Taking such a simple Graptolite as G. Sagittarius (fig. 27, i) 

 as the type of the sub-class, the hydrosoma is found to consist 

 of the " solid axis," the " com-mon canal," and the " cellules." 

 The entire polypary is corneous and flexible, and the solid 

 axis is a cylindrical fibrous rod, which gives support to the 

 entire organism, and is often prolonged beyond one or both 

 ends of the hydrosoma. The common canal is a tube which 

 encloses the coenosarc, and gives origin to a series of cellules, 

 these being little cups corresponding to " hydro thecae," and 

 enclosiftg the polypites. Not only are the essential details of 

 the structure — with the exception of the solid axis — strictly 

 comparable with that of a Sertularian, but there is a good 

 evidence, as shown by Hall and the author, that the reproduc- 

 tive prodess was al^o carried on iii a manner similar to what 

 we have seen in the other Hydroida — namely, by generative 

 buds or gbnophores. 



Wo Graptolite, however, has' hitherto been certainly proved 

 to have been fixed by a " hydrorhiza;" and it is only in 



