INFLUENCE OF MUSCULAR EXERTION UPON METABOLISM. 213 
A second experiment,* begun after fifteen hours’ fasting, was 
divided into two periods. The first was similar to the previous 
experiment, but lasted for thirty minutes only, the work of ascent 
equaling in that time about 30,000 kgms. The subject then rested 
for a time during which he consumed 105 grams of butter. Two 
hours after the ingestion of the butter the experiment was repeated, 
samples of the expired air being taken for three minutes at a time. 
The results as regards the respiratory quotient were as follows: 
Fasting. 
Three minutes before beginning work........ 0.706 
Twelfth to fifteenth minute................... 0.804 
Twenty-seventh to thirtieth minute.......... 0.812 
FRES bie sssaares tatty beadlard ence edlia dS icarias ata eed 0.812 
Two Hours after Ingestion of Butter. 
Three minutes before beginning work......... 0.666 
Twelfth to fifteenth minute................... 0.783 
Twenty-seventh to thirtieth minute.......... 0.809 
In conjunction with Laulanié + he has also experimented on 
dogs and rabbits, the muscular contractions being caused by electric 
shocks. The method of determining the respiratory exchange, as 
described by Laulanié, consisted in using a Pettenkofer type of 
apparatus with a small but constant known rate of ventilation. 
The outgoing air passed through a small gasometer, but the current 
could be shunted and the sample of air contained in the gasometer 
analyzed. No details of the experiments or of the methods of cal- 
culation are given. The first table on the following page contains 
Laulanié’s summary of the results. § 
An even greater increase in the respiratory quotient has been 
vbserved by other investigators. Thus Hanriot & Richet || found 
* Comptes rend., 122, 1169 
+ Ibid., 122, 1244, 1303: Archives de Physiol . 1896, p. 572. 
t Archives de physiol . 1896, pp. 619 and 636. 
§ Energetique Musculaire, p. 70. 
|| Comptes rend., 104, 435 and 1865; 105, 76. 
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