ORGANIZATION OF THE FOSSIL PLANTS OF THE COAL-MEASURES. O17 
in Calamostachys. As no peltate expansion is preserved in the specimens, there is 
no evidence as to the mode of attachment of the sporangia. 
In tangential sections the sporangia have an elongate, oblong form, thus differing 
from those of Calamostachys, which appear square in the corresponding view. This 
difference, however, seems to depend only on the oblique position of the sporangia in 
the “ Calamitean ” fructification. 
The sporangial wall resembled that of Calamostachys in structure. The spores are 
all of approximately equal size, and average ‘075 millim. in diameter. As sections 
were cut from all parts of several strobili, including their bases, it is not probable that 
macrosporangia, if they had existed, could have been missed. 
The anatomy of the peduncle is exactly that of a young stem of Calamuites, as above 
described. The pith is fistular, only the peripheral zone being persistent. The wedge- 
shaped bundles, sixteen to twenty in number, forma ring. Each bundle has a canal 
at its inner margin. We have been able to prove that here, as in the vegetative 
stems of Calamites, the canal contains the remnants of the protoxylem of the bundle. 
In longitudinal sections the spiral thickenings of the primitive tracheides in the canal 
can be easily seen.* 
Secondary wood, distinguished by the radial seriation of its elements, was formed 
in the peduncle just as in the vegetative stem. In fact the peduncle agrees in every 
respect with the typical vegetative axis of a young specimen of Calamites. 
The axis of the strobilus has essentially the same structure as that of the peduncle, 
of which it is the prolongation. The number of the bundles, which is equal to that of 
the sporangiophores, remains constant. Their structure is unaltered, except that, as 
we ascend the axis, we leave the secondary wood behind. At the nodes the 
arrangement of the tracheides is identical with that in Calamites. The bundles 
however show a slight difference in their arrangement. In the peduncle they are 
equidistant from one another; in the axis of the strobilus they become approximated 
in pairs. The cortex contains a ring of large cavities, half as numerous as the bundles, 
and alternating in position with the bundle-pairs. These cavities, which recall the 
“vallecular canals” of Eguisetum, ave best seen at the nodes, where the cortical 
tissues are best preserved. 
Now there can be no doubt that this remarkable fructification, specimens of which 
are unfortunately extremely rare, is that of a Calamites. If the peduncle were found 
alone it could not be distinguished from the stem, of corresponding age, of a typical 
Arthropitoid Calamite. . 
We wish to call attention to the very close resemblance between this strobilus of 
Calamites, and that described by M. Renautt under the name of Volkmannia gracilis, 
but placed by Count Sotms-LavpacH in Weiss’s genus Paleostachya.t In the 
* ©. N. 1569 and 1573. a 
+ Renavtt, ‘ Ann. Sci. Nat., Bot.,’ Sér. 6, vol. 3, Plate 2; also ‘ Cours de Bot. Fossile,’ vol. 2; Sous, 
‘ Fossil Botany,’ p. 832. 
