732 PROFESSOR W. C. WILLIAMSON AND DR. D. H. SCOTT ON THE 
2. The finer vascular bundles were probably collateral. 
3. There was a hypoderma towards the upper surface * 
5. Ona Peculiar Bud-like Structure. 
Before leaving the subject of the foliage, we wish to call attention to a very 
peculiar and indeed unique structure, which is represented in Plate 24, figs. 14 and 15. 
The organ at first sight strongly suggests the idea of a cone, and therefore attracted 
our special attention in the hope that we were at last on the track of the long-sought 
fructification of Lyginodendron. Subsequently, however, we found reason to give up 
this idea. The organ consists of an axis, imperfectly preserved in our specimen, 
which is densely clothed with prominent appendages, bearing a close resemblance to 
the well-known cortical emergences of the stem and leaf of Lyginodendron. The 
appendages are elongated and somewhat tapering; each has an envelope of scleren- 
chyma which becomes solid at the top. The central tissue is parenchymatous and 
has often perished, leaving a hollow. In no case is there any trace of a vascular 
bundle in an appendage. The transverse sections of the appendages are more or less 
semicircular near the base, becoming circular in their free part. 
The axis of the whole structure, where it is cut radially, is found to be hollow (see 
fig. 14). In one of the sections the surface of the axis is shown in tangential view, : 
and is seen to be marked with a number of patches of lighter coloured parenchyma, 
which no doubt represent the bases of as‘many appendages (see fig. 15). Between 
the parenchymatous areas the superficial tissues of the axis appear to have been 
fibrous, but the preservation is very imperfect. 
We have two sections of this curious object. The extreme length of the whole 
structure is 14 millims., but it is doubtful whether the whole of this belongs to one 
specimen. The continuous portion represented in figs. 14 and 15 is only 4 millims. 
long. 
We have rejected the idea of this specimen being a cone for two reasons :— 
1. The appendages show no trace of vascular tissue, and therefore could hardly 
have been either sporophylls or sporangiophores of any kind, while on the other hand 
they agree exactly with the cortical emergences of Lyginodendron. 
2, There is no trace of sporangia in connection with the organ. 
The evidence for the specimen belonging to Lyginodendron at all depends solely on 
the structure of the appendages, the identity of which, with the cortical outgrowths 
of that plant, can scarcely be doubted, if, for example, our figures 14 and 15 be 
compared with Memoir XVII, Plate 12, fig. 6. 
The explanation we would suggest is, that the structure represents either a bud or 
more probably a very young leaf covered with protective appendages of the nature 
* The best: preparations for showing the structure of the leaflets are: C.N. 1196, and others of the 
same series, 1856, 1885 D, and 1979. 
