620 OLIGOCHAETA 



in XIII ; pouches in IX- XI ; gizzards in five segments from XVIII. Ko 

 penial setae. No true spermathecae (?). Large spermathecal sac communi- 

 cating with egg-sacs and ovarian sacs, and giving oflF a tube passing round 

 gut fusing with its fellow of opposite side. 



This genus, like the last, contains only one species. As in Hyperiodrilus there 

 are paired penes, which are to some extent variable in position ; in one individual 

 they were upon the fourteenth, in another upon the fifteenth segment. A groove 

 starting from the male pore passes straight along the median ventral line of the body 

 dividing just at the penes into two branches which go to each penis. 



The testes are attached to the anterior wall of segments x, xi ; both are enclosed 

 in sacs, which also include the sperm-duct funnel ; the latter is, as in Hyperiodrilus 

 and several other genera, dilated before their termination in the funnel and traverse 

 the septum, from which the funnel depends, twice ; the funnel being thus situated 

 a segment behind that which it ought to occupy. I have, howeverj pointed out 

 in considering Hyperiodrilus that this difference from other worms may be more 

 apparent than real, since the cavity of the sperm-sac really belongs to the body 

 cavity of the segment in front of that in which it lies. 



The female organs of generation are rather different from the corresponding 

 organs of other genera. The ovaries are each enclosed in a sac which passes into 

 a tube soon dilating somewhat ; from this dilatation a narrow tube passes forward 

 and communicates with the long spermathecal sac ; this sac is divided into two parts 

 by a median constriction ; at this constriction the thickness of the lining epithelium 

 is much increased, so much so as to obliterate the lumen. There is thus apparently 

 no connexion between the narrower anterior section of the spermathecal sac and the 

 broader hinder part. Possibly, however, the arrangement is such that the sperm 

 can force its way in the one direction, but is unable to make way in the reverse 

 direction. It is also conceivable that the division of the sac corresponds to the 

 distinction between an epidermic and a mesoblastic portion of the pouch. From the 

 dilatation on the ovarian tube already referred to arises a widish tube which passes 

 round the gut and unites with its fellow on the opposite side. Another tube passes 

 into the egg-sac, with which, of course, is connected the oviduct; the latter, however, 

 also opens into the ' Eitrichterblase ' (= the dilated sac). The oviduct has the 

 peculiarity of possessing a diverticulum bound up in the same sheath with it ; with 

 this may be compared the diverticulum of the sperm-duct in Phreodrilus (cf. 

 •P- 273)- 



