XIVj LYCOPODIUM 43 



While several species possess a cortex of three distinct zones 

 (fig. 125, H, c, c', c"), in others the extra-stelar tissue is much 

 more homogeneous, consisting of thin-walled parenchyma or in 

 some cases of thick- walled elements ; as a general rule, however, 

 there is a tendency towards a more compact arrangement in the 

 inner and outer portions of the cortex as contrasted with the 

 larger and more loosely connected cells of the middle region. 

 In certain types the middle cortex contains fairly large spaces, 

 as in the swamp-species L. inundatum, which with L. alopecu- 

 roides exhibits another feature of some interest first described by 

 Hegelmaier ^ If a transverse section of the stem of L. inundatum 

 be examined the leaf-traces are seen to be accompanied by a 

 circular canal containing mucilage which extends intQ the lamina 

 of the leaf. In a specimen of L. cernuurn^ obtained at a height 

 of 2500 ft. by Professor Stanley Gardiner in the Fiji Islands, 

 the leaf- traces (fig. 125, B It,) were found to be accompanied for 

 part of their course by a well-marked secretory space (fig. 125, 

 B, s). There is little doubt that the presence of these mucilage 

 canals is directly connected with a certain type of habitat^ and 

 attention is called to them in View of a resemblance which they 

 offer to a characteristic strand of tissue, known as the parichnos, 

 which is associated with the leaf-traces of Lepidodendreae and 

 Sigillarieae. In the section shown in fig. 125, H, the xylem 

 of the stele forms more continuous bands than is often the case 

 in L. cernuum which has already been described as having its 

 xylem in small detached groups. The presence of the smaller 

 branch -stele (fig. 125, H, b) affords an example of monopodial 

 branching. The outer cortex of L. saururus (fig. 125, C) 

 exhibits a somewhat unusual feature in the distribution of the 

 thicker-walled tissue (b) which encloses a patch of more delicate 

 parenchyma (a) with large lacunae (Ic) in the region of the 



1 Hegelmaier (72). SeealsoHill,T.G.{06) p.269; this author draws attention 

 to the fact that in some species of Lycopodiam the mucilage canals are confined 

 to the sporophylls. 



2 Professor Yapp has drawn my attention to the very close anatomical 

 resemblance between a specimen of Lycopodium salakense obtained by him 

 from Gunong Inas in the Malay Peninsula and L. cernuum as represented 

 in fig. 125, H and I. 



8 Jones (05). 



