562 SYSTEMATIC SYNOPSIS. — COLUMBZ — PERISTERZE.’ 
seems best to draw the line, if one must be drawn, so as to include the Pterocletes in Columbe, 
and leave the Cracide and Megapodide with Galline. The Sand-grouse (better Sand-pigeons), 
or Péerocletes, represent the inosculation of the two series. They are terrestrial Columbines, 
modified for a grouse-like life; the digestive system is fowl-like (cceca several inches long, etc.); 
but the pterylosis, the sternum and humerus, the cranial and many other characters, are pigeon- 
like. The only alternative to reference of Pterocletes to the Columbine series is their elevation 
to independent ordinal rank, as proposed by Huxley. 
‘The Columba, as above indicated, are intended to be made conformable to Huxley’s Periste- 
romophe plus Pterocletes. Assuming the imperfectly-known extinct Dodo, Didus ineptus, to 
have been a modified Columbine, and considering the Pterocletes to represent a rasorial modi- 
fication of the Columbine series, the Order Columbe may be separated into THREE groups, or 
suborders, Dip1, PrEROCLETES, and PERISTER#, the first two certainly, the last probably, of_ 
a single family. The Peristere alone are American. 
9. SuBoRDER PERISTER: Trur CoLtumsBine Birps. 
(Equivalent to the Peristeromorphe of Huxley; the Gemitores of Macegillivray, or 
Columbe proper of most authors; the Gyrantes of Bonaparte, plus Didunculus; Columbe of 
Garrod minus Pterocletes; Pullastre of Liljeborg minus Cracide and Megapodide.) Skull 
schizognathous, schizorhinal; basipterygoids prominent; angle of mandible not produced; 
rostrum externally as above said. Sternum doubled-notched or notched and fenestrate, on 
each side; pectoral crest of humerus salient, acute. Carotids two. Syringeal muscles one 
pair. Cceca coli small or wanting; gizzard muscular; crop developed; gall-bladder generally 
absent. Fourth gluteal muscle undeveloped; second pectoral specially inserted; ambiens 
normally present, or wanting. Oil-gland nude, small, or wanting. Plumage without after- 
shafts. Feet insessorial; hallux normally insistent; tarsus normally seutellate. Rectrices 
normally 12 or 14. (Rasorial tendency in more rectrices, hallux up, and tarsus reticulate.) 
Altricial; psilopedic ; monogamous; eggs few. One family ? 
384. Family COLUMBIDZ: Pigeons. 
The family is here taken td be co-extensive with the 
suborder as defined. With one exception (Starnenas 
cyanocephala), all our species will be immediately recog- 
nized by their likeness to the familiar inmates of the 
dove-cot. One seemingly trivial cireumstance is so con- 
stant as to become a good clue to these birds: the frontal 
feathers do not form antize by extension on either side of 
the culmen, but sweep across the base of the bill with a 
strongly convex outline projected on the culmen, thence 
rapidly retreating to the commissural point. The plumule- 
less plumage is generally compact, with thickened, spongy 
thachis, the insertion of which will seem loose to one who 
skins a bird of this family. The head is remarkably 
small; the neck moderate; the body full, especially in the 
pectoral region. The wings are strong, generally length- 
ened and pointed, conferring a rapid; powerful, whistling 
flight ; the peculiar aérial evolutions that these birds are 
Fig. 389.— European Ring Dove(Co- wont to perform have furnished a synonym for the family, 
tumba palumbus). (From Dixon.) Gyrantes. The tail varies in shape, from square to grad- 
uate, but is never forked; as a rule there are 12 rectrices, frequently increased to 14, rarely to 
