36 DIVISION I.—GENERAL MORPHOLOGY. 
curved furrows ; and as gonidia are formed on the surface in a way that will be after- 
wards described (see Division II) the body may be termed a gonidiophore (Fig. 16). 
The hyphae of the Fungus-body must necessarily make their way for some distance 
from the ovarian base into the floral pedicel, for it is difficult to conceive of any other 
mode of supplying food to the Fungus; but we have no exact information on this 
point. When the gonidiophore is fully formed, the beginning of a sclerotium makes 
its appearance in the torus at its base and on the mycelium, which is supposed to 
spread through it, in the form of a small somewhat elongated fungal body enclosed 
in the white tissue and distinguished by its greater density (Fig. 16 4, s). It is 
formed at first of slender delicate separable hyphae which are continuous with 
those in the vicinity, but are somewhat firmer and more closely compacted. Its 

Fic. 16. Claviceps purpurea, Tul. a young ovary of FIG. 17. Claviceps purpurea, Tul., on Secale cereale. 
Secale cereale penetrated and covered with the gonidio- a seen from without. 4 median longitudinal section. The 
phore, seen from without; the hairs of the ovary and the sclerotium s rests on the torus and carries up the dry- 
remains of the style g project at the apex from the fungal ing gonidiophore ? on its apex. After Tulasne. Slightly 
inve é longitudinal section gh a similar stage magnified, 

in the development from Seca/e; s commencement of 
the sclerotium. c¢ similarly young state of the Fungus on 
the .pistil of Glycerta fluttans, the Fungus projecting 
beyond the apex of the ovary. After Tulasne. Slightly 
magnified. 
surface soon acquires a violet hue, the superficial cells beginning to assume the 
character of the future rind. It now increases in thickness and elongates into the 
well-known horn-shaped body, which is attached at its base to the torus and projects 
above from between the paleae. The course of its development still requires more 
exact investigation. Its growth in the longitudinal direction is no doubt maintained 
by continued addition at the base. The increase in thickness of each transverse 
section above the base must in a great measure be due to the expansion of cells 
already formed, since these are more than four times broader in the fully 
developed parts than in the younger. The gonidiophore ceases to grow as soon as 
the sclerotium begins to be formed, and being detached from the torus as the 
sclerotium enlarges it is carried up like a cap on its apex, and there shrivels up and 
sooner or later falls off (Fig. 17). 
