CHAPTER III.—SPORES OF FUNGI. 99 
again from their mother-cells, such as asci and the like. Initial cells of the same 
morphological value may in case A become spores directly, in case B may become by 
division aggregates of spores, which were named above compound spores, but which 
it would be better perhaps to designate by- the term previously suggested of sfore- 
groups or sporiderms. The descriptions would thus be rendered short and clear, and 
would stand the scrutiny of the attentive and thoughtful student. The result would justify 
the experiment. 
That compound spores are produced by ordinary bipartition with septation of their 
initial cells, as represented in Fig. 49, has been recently confirmed by Strasburger'. I do 
not here go further into the subject of Körber’s sporoblasts and the correlative terms”. 
The long filiform initial cells of species of Cordyceps® divide into a very large number 
of short cylindrical simple spores, but it is still a question whether the division is 
simultaneous or by successive bipartitions. In these Fungi moreover the compound 

FIG. 49. Sphaeria Scirpi, DC. a—« jive stages of devel of the spores in the order of the letters; 
all drawn from specimens inside recently isolated and uninjured asci. /ripe ip d spores disch from the 
ascus, After Pringsheim. a—e magn. 390, / 350 times. 


spores, eight of which are formed in each ascus, break up of their own accord when 
they escape into their numerous component parts, each of which can germinate by itself; 
thus they are disuniting spore-groups, and their disunion may take place inside the 
ascus. .We should be obliged to say in this case, according to the usual terminology, 
that each spore breaks up into many spores capable of germination. In nearly related 
genera, such as Claviceps, the filiform spores remain undivided till after germination, 
and put out germ-tubes. The spores also of Cenangium fuliginosum, Fr. behave in 
the same way as those of Cordyceps, while the spores of other species of Cenangium 
do not disunite*. De Notaris® gives an account of the disunion of the spores of 
Sporormia fimetaria, Not. 
II. STRUCTURE OF THE RIPE SPORES. 
Section XXX. In considering the structure of the ripe spore we must 
distinguish between moszonless spores and those endowed with power of motion or 
swarm-spores. 
The motionless spores are much the most common, and everything that was 
said in previous sections, with the exception of sections XVIII. aand XX. a, about the 
development and discharge of spores, referred only to them. In the mature state they 
are cells of extreme variety of form in the different species, most frequently round 
or longer than broad, in Claviceps and some others elongated cylindrical tubes. 

1 Zellbild. u. Zelltheil. ed. 3, p. 51. 
2 Syst. Lichen. German. Einleitung. 
® Tulasne, Carpol. III; Bot. Ztg. 1867, 1. 
* See Tulasne in Ann. d. sc. nat. XX, ser. 3, p. 135. 
5 Microm. Ital. Dec. V in Mem.R. Acad. d. Torino. 
H 2 
