140 DIVISION II,—-COURSE OF DEVELOPMENT OF FUNGI. 
thorough investigation; Pringsheim’s Pythium endophytum! and Schenk’s Achlyo- 
geton? are certainly very near it. 
Literature of the Ancylisteae. 
CORNU, Monogr. des Saprolegniées in Ann. d. sc. nat. ser. 5, XV, and im Bull. soc. bot. 
de France, XVI (1869), p. 222 ;—-Id., cited in Sachs, Traité de Bot. translated by 
Van Tieghem, p. 328.—See aiso Schenk, Ueber contractile Zellen, Wiirzb. 1858, p. 9. 
ZoPF in Bot. Ztg. 1879, p- 351. 
PFITZER in Monatsber. d. Berlin. Acad. 1872, p. 351. 
MONOBLEPHARIS. 
Section XXXIX. The aquatic genus Monoblepharis with three species, which 
at present has been examined only by Cornu’, is related to the Peronosporeae. 
These plants, according to Cornu’s short and somewhat incomplete description, 
resemble the Pythieae in their vegetative structure and in their mode of life. They 
form on their thallus sporangia with swarm-spores, and the latter originate and 
escape, not in the manner of 
Pythium, but in that of Phyto- 
phthora, &c. and have only one 
cilium. Oogonia and anther- 
idia are terminal or intercalary 
on the branches of the thallus ; 
their disposition, which varies 
in the different species, agrees 
with that of some species of 
Pythium ; Fig. 67 shows it in 
Monoblepharis sphzrica. The 
points in which M. differs from 
Pythium appear in its further 
development. Firstly, the whole 
FIG. 67. Monoblepharis sphaerica, extremity of afi bearing an ium Of the protoplasm of the oogo- 
oand an antheridium a. x. before the formation of the oosphere and spermatozoids. . j A = 
2 ns rn which is an ee ven van Krisen 
After Cornu. Magn. 800 times. globules is transformed into the 
oosphere without separation of 
periplasm and with diminution of volume, and as this takes place the wall of the 
oogonium opens at its upper end. Secondly, a few swarm-cells (spermatozoids) 
are formed by division of the protoplasm in the antheridium; the spermatozoids 
escaping through an aperture in the wall of the antheridium move with a gliding 
motion over the wall of the oogonium, till one of them finds its way through the 
aperture to the oosphere and coalesces with it. The oosphere is thus fertilised and 
an investing membrane is formed, which subsequently becomes strongly thickened 
and rough with warts on the outer surface; in this state the body is a resting oospore, 
the further development of which is unknown, but can hardly differ from that of the 
Peronosporeae. 





1 Jahrb. f. wiss. Bot. I. ? Bot. Ztg. 1859, p. 398. 
® Ann. d. sc. nat. ser. 5, XV, 1872. 
