294 DIVISION II.—COURSE OF DEVELOPMENT OF FUNGI. 
containing tissue lies originally between each pair of future lame] ae, which does not , 
however grow with them, but disappears as the general structure developes and thus 
completes the delimitation of the surfaces of the lamellae (Fig. 135 g). The dispute 
is not important and arises perhaps from specific differences. But there is no 
question that the primordia of the lamellae have no free lateral margin; their 
longitudinal margin on the side of the stipe passes into the tissue of the surface 
of the stipe as continuously as the margin that looks towards the pileus passes into 
the substance of the pileus. 
The commencement of all the principal parts is followed by a general vigorous 
growth throughout, and from this time the pileus and the stipe grow more rapidly 
than the bulbus, the contrary having been the case in former stages. The portion of 
the stipe above the bulbus, hitherto a Jow narrow disk surrounded by the commencing 
hymenium, grows into a stout body slightly conically pointed at the apex, and the 
growth of the young pileus and of the hymenium keeps pace so exactly, that the 
attachment of the lamellae to the surface of the stipe, which has been already noticed, 
is not interfered with; the pileus increases in size by superficial extension in the 
direction of the margin, and the lamellae which are first formed increase 
correspondingly in radial length in the same direction, and in breadth in the 
direction of the stipe. The increase in breadth, however, is comparatively small, so 
that the marginal portion of the pileus is somewhat curved over and bent downwards 
(epinastically) in towards the surface of the stipe (Fig. 135 d). Lastly, as the general 
surface of the hymenium becomes broader towards the margin in consequence of 
this growth, new lamellae make their appearance between the original ones, being 
distinguished from them only by the fact. that the edge towards the apex of the 
pileus does not extend as far as the apex, while that towards the margin lengthens as 
the margin advances ; the later they are formed the greater the distance from the apex 
of the pileus at which they terminate. All this growth of pileus and lamellae 
marginwards takes the same course as regards its direction as in the rest of the 
Agaricineae ; but it does not result in the formation of free margins or even of 
margins bounded by gaps. The various margins are bounded by undifferentiated 
hyphal weft with intercalary growth and pass into it without any distinct boundary 
line, and at the zone of transition the differentiation is constantly spreading over 
newly formed sections of this weft. 
These processes of growth and differentiation at length reach their limit, and 
are followed by the final completion of all the parts, the last acropetal elongation of 
the now tall cylindrical stipe and the hyponastic upward expansion of the pileus 
—phenomena which again follow in their main features the same course as in the 
species previously described. There is only one more peculiarity to be observed in 
the Amaniteae, namely, that the peripheral layer of the stipe into which the edges of 
the lamellae pass, a thick loosely felted layer, takes no part in the elongation of the 
inner portion of the stipe which it encloses. It separates everywhere as the 
elongation proceeds from the surface of the stipe, retaining its connection with it 
only at the point of insertion of the pileus (Fig. 135 d); it also continues at first in 
connection with the edges of the lamellae, forming a continuous membrane which 
extends over the whole surface of the hymenium up to the margin of the pileus. As 
the pileus expands the connection is dissolved, beginning from the margin of the 
