62 INTESTINAL MEASUREMENTS 



structure. It is held by Gegenliaur that this organ is the 

 equivalent of the reptilian tongue, a,ud that in the skeletal 

 vestiges which it contains are to be found the equivalents of the 

 hyoid skeletal cartilages wliich support the tongue in lizards. 

 In this case the tongue of mammals is a subseijuently added 

 structure. 



The oesophagus leads from the mouth cavity to the stomach. 

 The latter organ has commonly a distinctive shape in mammals. 

 This is well shown in Alan. The uritices of the oesophagus and 

 intestine are somewhat approximated ; and this causes a bulging 

 of the lower border of the organ, usually spoken of as the greater 

 curvature. A stomach of this typical form is found in many 

 orders of mainmals, and is unlike the stomach in any of the 

 groups of lower vertebrates in shape. Sometimes the shape of 

 the organ is greatly altered : it may l^e dia\Mi out, sacculated, 

 or divided, as in the Euminants and Whales, into a series of 

 differentiated chambers, each of wliich plays sonre special part in 

 the phenomena of digestion. 



The intestine of mammals is always long and much coiled, 

 though the length and consequent degree of coiling naturally 

 varies. On the whole it is perhaps safe to say tlia't it is shorter 

 in carnivorous than in ^-egetable-feeding beasts. Thus the Paca 

 has an intestine of ;!1) inches total length, while the Cat, an 

 animal of about the same size, has an intestine which is only 

 36 inches long. .V fish diet, however, to judge from the Seals, 

 is associated with a long intestinal tract. The intestine is 

 di\isible in the vast majority of mammals int(j a small and a 

 large intestine. The two are separated by a valvular constriction 

 save in certain Carnivores ; and in the majority of cases the 

 distinction is als(.) ciiiphasised by the ijresence at the junction of 

 a blindly-ending diverticulum, tlic ciiecum. This latter organ 

 varies greatly in length, being very short in the Cat-tribe and 

 exceedhigly long in L'odents. Its size is, to some extent, de- 

 pendent upon the flesh -eating or grass -eating propensities of 

 the animal in which it occurs, ihiv. of the longest caeca is 

 possessed by the Vulpine Phalanger, in whicli the organ is one- 

 fifth of tlie length of the small intestine ; while the opposite 

 extremity is reached l.)y Felis man-Dscelis, which has a small 

 intestine one hundred times the length of the caecum. 



Aw interesting ])oint in connexion with the gut of mammals 



