CNOTHERA AND THE MUTATION THEORY 309 
Frequency of appearance from different races 
New or “mutant” ~ 
ae Ta ick lata oo. simplex | nanella | oblonga} gigas 
oblonga........ 0.7 0.7 6.1 0.0 O02! [is ee soece 0.0 
nanella........ 0.5 0.4] 0.1 ee: || eee els 0.0 0.9 
lata. ........., Ov4 Psa 0.4 0.3 0.02 0.0 0.0 
scintillans..... 0.3 On. esehceos iy 0.3 0.0 0.0 0.0 
albida......... 0.2 2.1 0.1 0.0 0.0 0.1 0.0 
rubrinervis. .... 0.04 | 0.2] 0.0 0.0 0.0 0.2 0.0 
Ovata......... 0.01 0.4 0.0 0.0 0.0 0.0 0.0 
deserens....... 0.0 0.0 | 0.0 3.2 0.0 0.0 0.0 
metallica....... 0.0 0.0 0.0 1.5 0.0 0.0 0.0 
Total new types| 2.2 4.1] 6.7 6.5 0.05 | 0.3 0.9 
as early as 1902, to suggest that O. Lamarckiana 
might be a hybrid, and that the new types might 
be simply recombination products. There were 
very serious difficulties in the way of this hypothesis, 
and it was not strongly urged by its proponents. 
The view was, however, strengthened by a study of 
the history of the species. The members of the 
group to which it belongs are apparently all native 
in America, but several of them have become 
naturalized in Europe. It is very doubtful if 
O. Lamarckiana occurs in America as a wild plant, 
so that it may well be suspected of being a hybrid 
that was first obtained artificially and that then 
accidentally escaped and became established in 
Europe. Davis has worked on this hypothesis and 
has tried to synthesize the species by crossing wild 
American species. The question has now lost 
much of its point, however, through the discovery 
