136 Experimental Zoology 
These and other experiments with pigeons show that the results 
are more complicated than in the case of fowls. It is not possi- 
ble to bring the results under a single point of view at present. 
Ewart’s experiments’ with pigeons indicate that while certain 
kinds of crosses may give rise to offspring resembling the Rock 
Pigeon, yet in other cases a more immediate ancestral color may 
comeout. A dark blue Fantail, having all the characteristic bars 
of the Rock Pigeon, was bred to a less pure blue Fantail. On 
two occasions an absolutely pure white Fantail was produced. 
This result, Ewart thinks, is due to a reversion to a white grand- 
parent. Ewart also crossed a white Fantail with a white Pouter. 
The offspring was white in color, but in form resembled the 
Pouter. A hybrid between an “archangel” and an “owl” was 
bred to a white Fantail. The two offspring were blue, one of 
them being almost identical with the wild Rock Pigeon, more es- 
pecially with the Indian variety. Not only was there reversion 
in color, but in form as well. 
Ewart seems to think that reversion amongst closely inbred 
races of dogs, horses, and pigeons leads to a sort of rejuvenes- 
cence of the stock. Those individuals showing the ancestral 
characters prove to be stronger and more active. 
The reversion to the type of the Rock Pigeon, that seems to play 
often so conspicuous a réle in these experiments with pigeons, 
recalls the return to the gray color in mice when fancy breeds 
are crossed, but in the second generation of mice there is a re- 
turn in some of the forms to the parent types. How far this 
occurs in pigeons is not clear from the evidence at hand. 
A brief but important note in alternate inheritance in pigeons 
is given by Staples-Browne. Webfoot sometimes suddenly 
appears in pigeons. A pigeon of this sort crossed with another 
having normal feet produced six normal-footed offspring. These 
individuals (F,), inbred, produced in one case nine with normal 
feet and three with webbed feet. Another pair (Ff), however, 
produced seventeen normal birds. Extracted web-footed 
individuals produced six web-footed. It appears that the latter 
condition is recessive to normal feet. 
1 The Penycuik Experiments. 
