272 MOVEMENTS OF THE POLLINIA, Chap. IX. 



is seen from above highly magnified in the upper of 

 the two adjoining figures ; and in the lower figure we 

 have a longitudinal section of the uncontraeted disc, 

 together with the base of the attached and upright 

 caudicla At the broad end of the disc there is a deep 

 crescent-shaped depression, bordered by a slight ridge 

 formed of longitudinally elon- 

 '^' ' gated cells. The end of the 



caudicle is attached to the 

 steep sides of this depression 

 and ridge. When the disc is 

 exposed to the air for about 

 thirty seconds, the ridge con- 

 tracts and sinks flat down ; in 



Disc of Gymnadenia conopsea. • i • -j. j -ii. -j. ±t. 



^ Sinking, it drags with it the 

 caudicle, which then lies parallel to the elongated 

 tapering part of the disc. If placed in water the 

 ridge rises, re-elevating the caudicle, and when re- 

 exposed to the air it sinks again, but each time with 

 somewhat enfeebled power. During each sinking and 

 rising of the caudicle, the whole pollinium is of course 

 depressed and elevated. 



That the power of movement lies exclusively in the 

 surface of the disc is well shown in the case of the 

 saddle-shaped disc of Orchis pyramidcdis ; for whilst it 

 was held under water I removed the attached caudicles 

 and the layer of viscid matter from the inferior surface, 

 and immediately that the disc was exposed to the air 

 the proper contraction ensued. The disc is formed of 

 several layers of minute cells, which are best seen in 

 specimens that have been kept in spirits of wine, 

 for their contents are thus rendered more opaque. The 

 cells in the flaps of the saddle are a little elongated. 

 As long as the saddle is kept damp, its upper surface 

 is nearly flat, but when exposed to the air (see fig. 3, 



