GENUS 37. THISTLE FAMILY. 443 
37. DOELLINGERIA Nees, Gen. & Sp. Ast. 177. 1832. 
Perennial leafy herbs, the lower leaves reduced to scales or sheaths, the upper large, 
mostly entire, acute or acuminate in our species. Basal leaves none. Heads corymbose, 
usually numerous; involucre campanulate to hemispheric, its bracts imbricated in several 
series, appressed, thin, sometimes scarious-margined, their tips not herbaceous nor foliaceous. 
Receptacle foveolate. Ray-flowers white, pistillate, not very numerous. Disk-flowers per- 
fect, the corolla with a slender tube abruptly expanded into a campanulate 5-lobed limb, 
white to greenish in our species. Anthers obtuse at the base; style-appendages ovate to subu- 
late (rarely obtuse). Achenes obovoid, glabrous, or pubescent. Pappus double, the outer 
series of numerous short bristles or scales, the inner series of long capillary bristles, some 
or all of which have thickened tips. [In honor of Th. Dollinger, botanical explorer.] 
About 5 species, natives of eastern North America. Type species: Doellingeria umbellata 
(Mill.) Nees. 
Leaves lanceolate to ovate; heads mostly numerous. 
Leaves lanceolate to oblong-lanceolate, acuminate. 1. D. umbellata. 
Leaves ovate to ovate-lanceolate, acute. 2. D. humilis. 
Leaves, at least the lower, obovate; heads commonly few. 3. D.infirma. 
1. Doellingeria umbellata (Mill.) Nees. Tall Flat-top White Aster. Fig. 4380. 
Aster umbellatus Mill. Gard. Dict. Ed. 8, No. 22. 1768. 
Diplopappus umbellatus Hook. Fl. Bor. Am.2: 22. 1834. 
D. umbellata Nees, Gen. & Sp. Ast. 178. 1832. 
Aster umbellatus var. pubens A. Gray, Syn. Fl. 1: Part 
2,197. 1884. 
D. pubens Rydb. Bull. Torr. Club 37: 147. 1910. 
Stem glabrous or pubescent above, striate, corym- 
bosely branched at the summit, 1°-8° high. Leaves 
lanceolate to oblong-lanceolaté, ascending, glabrous 
above, usually pubescent at least on the veins be- 
neath, membranous, acuminate at the apex, nar- 
rowed at the base into short petioles, or the upper- 
most sessile, hispid-margined, those of the stem 
5-6’ long, 4’-1’ wide; heads numerous, 6-10” broad, 
in terminal compound corymbs; involucre broadly 
campanulate or hemispheric, 14’-2” high, its bracts 
lanceolate, usually pubescent or ciliate, acutish or 
obtuse, imbricated in 3 or 4 series, the outer shorter ; 
rays 10-15, white; style-appendages ovate, acute; 
pappus nearly white; achenes nerved, slightly pu- 
bescent. 
In moist soil, Newfoundland to Georgia, Saskatche- 
wan, Iowa and Michigan. July—Oct. 
2. Doellingeria himilis ( Willd.) Britton. 
Broad-leaved Flat-top White Aster. 
Fig. 4381. 
Aster humilis Willd. Sp. Pl. 3: 2038. 1804. 
D. amygdalina Nees, Gen. & 25, Ast. 179. 1832. 
Aster umbellatus var. latifolius A. Gray, Syn. Fl. 1: 
Part 2, 197. 1884. 
Doellingeria humilis Britton, in Britt. & Brown, Ill. Fl. 
3: 392. 1898. 
Gi 
nay eM 
WY 
a Similar to the preceding species, usually lower, 
seldom over 4° high; stem striate, corymbosely 
branched above, glabrous, or somewhat pubescent. 
Leaves ovate to ovate-lanceolate, rather firm, ses- 
sile, or the lower very short-petioled, acute or short- 
acuminate at the apex, narrowed, or sometimes 
rounded at the base, rough-margined, those of the 
stem 1’-3’ long, 4’-2’ wide, inflorescence as in D. 
umbellatus; pappus dirty white; achenes somewhat 
pubescent. 
In moist: soil, eastern Massachusetts to New Jersey, 
Pennsylvania, Florida and Texas. Perhaps a broad-leaved race of the preceding species, July—Sept. 
