= 409 me 
brought together for examination, including specimens so well pre- 
served as to admit the delineation of the epidermal cells and stomata, 
no recognized fern type of fructification has yet been discovered. This 
evidence for a cycadofilicate nature in the genus is, of course, purely 
negative. 
On the basis of the characters of the sterile fronds the genus has 
generally been classified with the Zaeniopterideae or the Dictyopteri- 
deae, which are composed chiefly of types now placed in the Pteri- 
dosperms (Cycadofilices). In an-earlier paper. (1) I have suggested the 
genetic relationship of Glossopteris to the Megalopterid group in which 
I placed Yaeniopteris, Neuropteris and Lesleya. (2) The genus las 
named, from the base of the Upper Carboniferous, may well have been 
ancestral to Glossopteris, from which it differs only by the absence of 
anastomosis in the nervation. 
The fragment described below as ‘‘Glossopteris sp.,’’ and illus- 
trated in Pl. vil, Figs. 5 & 6. almost certainly represents sporangi- 
ferous organs of Glossopteris comparable to the cycad inflorescence. 
For the reasons just referred to I place both Glossopteris and its 
apparently near relative Gangamopteris in the Pteridosperms, not- 
withstanding the fern-like character of the rhizome Vertebraria. To the 
same groupe is to be referred, as I am convinced, the sporangiferous 
organs described as Ottokaria, and Derbyella. . 
Glossopteris Browntana Brongn. 
Pl. vi, Figs. 3, 4, 6 
1828. Glossopteris Browniana Brongniart, Prodrome, p. 
54 (nomen nudum); Goeppert, Syst. fil toss. 1836, p. 346, pl. 
xxi, fig. 9, 10; Morris in Strzelecki, New South Wales, 1845 p, 
247, pl, vi, figs. 1, 1a; Dana, Wilkes Expl. Exped., vol. x, 
1849, p. 716, pl. xxii. figs. 18a, 13b, 13c ; Tate, Quart, Jour. 
Geol. Soc., vol, xxiii, 1867, p. 140, pl. vi, figs. 5, 7; Feistmantel, 
Palaeontogr., Suppl]. IIT, 1878, p. 78, pl. viii, figs, 3, 4, pl. x, 
figs. 1, 3, 4, 5, 7, pl. xi. fig. 4, Feistmantel, Fl. Gondwana 
Syst., vol. iii, pt. 2, 1880, p. 102, pl. xxviA, fig. 2, pl. xxviiA, 
fig’. ©, pl, NSIKA, gs, 1, 2,9, 6, 8. Pl. SIA, Tg. 3; Feistmantel. 
(1) Bull. Geol. Soc. Amer., vol. TV, 1893, p. 128. 
(2) See Lesquereux: Goal Flora, vol. 1, p. 143, pl. xxv, fig. 1-3, 
