(/ YMNOSPERM^. 



403 



iogue of a macrospore, which here is not freed from the 

 parent plant. The endosperm clearly bears the same rela- 

 tion to the embryo sac as the prothallium of Isoetes does to 

 the -macrospore f and the corpuscula are slightly modified 

 archegonia. In some corpuscula the resemblance to arche- 

 gonia is very marked, the germ-cell below being surmounted 

 by a short neck ; Strasburger has even discovered a rudi- 

 mentary axial-cell, thus completing the correspondence of 

 these organs to those of the 

 higher Pteridophytes. 



512; — Fertilization is eflect- 

 ed by means of the pollen, 

 which comes in contact with 

 the apex of the ovule. It is 

 transported from the male 

 flowers mostly by the wind, 

 which accounts for the im- 

 mense quantity produced. 

 When the ovule has reached the 

 proper stage the micropyle is 

 filled with a fluid, which, dry- 

 ing, cai'ries the adherent pollen 

 grains into contact with the 

 apex of the ovule body, where 

 they germinate and form pol- 

 len tubes ; the latter penetrate puscuium ; p, two poiien grains ap- 



,T ni J • J. Ji 1 T plied to the apex of the ovule body, 



the soft tissue OI the ovule and fnto which they have sent two pollen 



eventually reach the corpus- tubes, «, ».-After p,anti. 

 cula (Fig. 299). In those cases where the corpuscula are 

 separated from one another each pollen tube comes in con- 

 tact with only one corpusculum (Figs. 397, B, and 399) ; but 

 when the corpuscula are close together a single pollen tube 

 may come in contact with all of them (Fig. 398, 1 and 2). 

 The union of the protoplasm of the pollen tube with that of 

 the germ-cell appears to take place by diffusion through the 

 wall of the former, as no openings in it have been discovered. 

 After fertilization the protoplasm in the germ-cell becomes 

 more turbid and granulnr, and soon at the base a transverse 

 partition is formed, cutting off a cell, which is the rudiment 



Fig. 399.— Diagrammatic i-cction of 

 an ovule of Pinvs, showing fertiliza- 

 tion, i, integument or coat of the 

 ovule : OT, the micropyle ; i, the body 

 or " nucleus" of the ovule ; e, the em- 

 bryo pac, filled with endosperm ; c. c, 

 two corpnscula shown filled with pro- 

 toplasm ; A, the neck cell of one cor- 



