CALAMOPITYEAE 479 



dermal fibres, as in Heterangium and other genera (Fig. 

 176, o.c). 



The cortex is traversed by the large leaf-trace 

 bundles (/./). The trace, when it first leaves the 

 wood, is a single bundle, but immediately divides into 

 two, precisely as in Lyginodendron. The leaves were 

 spirally arranged, and the phyllotaxis appears to have 

 followed the |- arrangement. Each transverse section 

 constantly includes the trace-bundles of five successive 

 leaves, which appear to have been more crowded than 

 in Lyginodendron, for the same transverse section may 

 show two leaf-bases in connection with the stem. The 

 petioles were of large size, even exceeding the stem 

 itself in diameter. 



The characters shortly described, according to the 

 investigations of Count Solms, to whom our whole 

 knowledge of the anatomy of this species is due, agree 

 so closely with those of Lyginodendron, that even a 

 generic separation would scarcely seem justified, if it 

 were not for two points which are still to be mentioned. 

 One of these is the fact that the leaf-trace bundles 

 appear to have assumed concentric structure immediately 

 on entering the cortex of the stem ; the other and more 

 important point is that, on entering tjie leaf-base, the 

 two bundles underwent further divisions, so that the 

 petiole contained half a dozen bundles or more, instead 

 of two at most, as in Lyginodendron. The leaf-stalks, 

 characterised by the ring of large vascular bundles and 

 the hypodermal strands of sclerenchyma, would fall into 

 the old genus Kalymma, one of the numerous fossil 

 genera founded on isolated organs of plants, which 

 subsequent research is gradually enabling us to identify. 



