222 GENETICS IN RELATION TO AGRICULTURE 
In one F» population of 255 plants Lotsy was able to distinguish about 
twenty-five different flower types as shown in Fig. 98. The flower 
types were not distinct, but represented merely different steps in an 
almost continuous series, save for the discontinuity incident upon the 
sharp segregation of a group of plants which bore peloric flowers. More- 
over, within any of these flower types the plants differed greatly in a 
number of other characters, such as size, color of flower, form of leaf, 
habit of growth, etc. As regards fertility there was segregation into 
self-fertile and self-sterile plants, the former being in the majority. Of 
the 255 plants, 185 produced zygomorphic flowers, 119 peloric flowers, 
and one plant produced both zygomorphic and peloric flowers. In 
color the flowers on different plants ranged from the deep red of the majus 
parent to the pale color of molle. 
Lotsy also grew several F3 populations. One of these from an Fy, 
plant bearing hooded zygomorphic flowers consisted of 209 plants all 
of which were different, indicating again an extreme condition of hetero- 
zygosity. Not a single plant produced flowers displaying the hooded 
character of the parent plant. There was again a vast array of flower 
forms, twenty-three different types being represented. With respect 
to the peloric condition, 113 plants bore peloric flowers only, 94 zygo- 
morphic flowers, and 2 bore both types. Although several different 
colors were represented, Lotsy was able to arrange them in two classes; 
the first consisting of 153 plants approximating the red color of majus, 
and the second group of 56 plants of about the color of the pale molle 
parent. There was, therefore, a fair indication of Mendelian segrega- 
tion for color in this generation. 
In this population as shown in Fig. 99 a plant was obtained which 
very closely resembled the type of A. molle in all its characters, and re- 
produced these characters in its progeny. Other plants were obtained 
which strongly resembled majus in certain of their characters, but not so 
completely throughout. The important feature here is the fact that 
even in F, segregation and recombination of factors have produced a 
plant which is practically identical with one of the parents. 
Other F’; populations were grown from F, plants displaying different 
sets of characters. One of these from a zygomorphic F» plant produced 
a population segregating for color and form (zygomorphic vs. peloric) of 
flower. Another population from a peloric F; plant consisted entirely of 
peloric flowering plants, but in this population there were many different 
color classes. An F2 plant of the pale color of molle gave an F'; popula- 
tion consisting entirely of pale-flowering plants but showing segregation 
in form and for the peloric character. 
Obviously if results such as these are to be explained on a Mendelian 
basis, it must be assumed that a relatively large number of factor differ- 
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