278 LEPIDONOTUS SQITAMATITS. 



Feet. — The second foot has a small dorsal division, which projects anteriorly, and no 

 dorsal cirrus. The dorsal bristles are tapering and spinous from the skin outwards, as in 

 other parts of the body. On the other hand, the ventral division has more slender bristles, 

 with longer and more tapered spinous tips than elsewhere. When seen antero-posteriorly 

 they somewhat approach the pinnately spiked forms of the Aphroditidas, though, of 

 course, differing in character. The long tentacle-like ventral cirrus comes from the 

 anterior and inner base of the foot, and is directed forwards, thus diverging both in 

 size and relation to the parts of the foot from the subsequent organs. 



The third has become more or less normal in structure, except that the ventral 

 division has not attained the size of that in the fully-developed foot. The serrated dorsal 

 bristles spring from an elevation on the upper and anterior border, the only representative 

 of the dorsal division, if we except the elevated crest bearing the dorsal cirrus. The 

 ventral bristles are stronger, the serrated tips somewhat shorter, and the points slightly 

 hooked. The slender inferior series alone still resemble those in the second foot. The 

 ventral cirrus has now the normal position and form. 



The fourth foot, though smaller, is nearly typical, and the short curved dorsal 

 bristles are also present. The ventral bristles have tips only a little longer than 

 those of the central feet. The ventral cirrus is in the normal position, but the long 

 papilla of the segmental organ does not appear till the eighth foot, and it continues to 

 the penultimate one. 



The typical foot (Plate XL1I, fig. 25) is massive, and presents little differentiation 

 of dorsal and ventral divisions other than the bristles and spines, for the main mass of 

 the foot is formed by the ventral division. The dorsal bristles rise from a papilla placed 

 far back on the dorsal edge and in the front of the cirrus, where that is present, and 

 their tips, as a rule, extend little beyond the bases of the ventral bristles. In structure 

 they consist of long, tapering, spinous bristles (Plate XXXVII, fig. 13), the spines being 

 arranged in close rows from base to apex, a few on the dorsal edge being shorter and 

 stouter, with coarser spikes (Plate XXXVII, fig. 12). The ventral bristles (Plate 

 XXXVII, figs. 9 and 10) have slightly longer tips than in L. clava, and the two spines at 

 the tip of the rows are characteristic. 



The shape of the foot does not essentially alter posteriorly, and the ventral bristles 

 of the last foot have tips only a little longer than in the centre of the body, a feature which 

 almost disappears in that next in front. 



The dorsal cirri spring from a dilated base (ceratophore) which has anteriorly and 

 posteriorly an area in which a whitish granular amorphous substance is deposited. 

 The deposit, indeed, marks the bulbous region on each side. The column is pale, but at 

 the dilatation is a blackish band, while the filiform tip is pale. 



The dorsal bristles are generally covered with fine mud and parasitic growths. The 

 general hue of the ventral is golden, but it varies in intensity, some of the northern 

 forms, e. g. from Norway and the Gulf of St. Lawrence, having darker bristles. In one 

 large example from St. Abb's Head one side had golden bristles, while in the other all were 

 of a pale colour. The same form had an enigmatical, hyaline, stalked structure, with 

 numerous rootlets on a posterior scale. An occasional foraminifer and a small patch of 

 Lepralia also are met with in large examples. 



