FOOTMARKS IN THE SANDSTONES OF CUMMINGSTONE 109: 
was a useful warning, however, to find that I should have been un- 
supported by the event, had I done so; for all the other remains 
depart, more or less widely, from the ordinary Crocodilian type of 
organization. The smallest amount of difference is perceptible in 
the femur, an incomplete cast of part of the shaft and distal end of 
which bone (of the left side) shows that it was thicker and stouter 
in proportion than that of the Crocodile. The impressions of the 
articular surfaces of the condyles, again, are so rough and irregular 
as to lead to the suspicion that they were covered by imper- 
fectly anchylosed epiphyses—which is the reverse of a Crocodilian 
character. 
The cast of the only example of a metacarpal or metatarsal bone 
which has come to light also exhibits proportions which indicate a 
much shorter and thicker foot than the corresponding bone in the 
modern Crocodiles, and, @ fortior7, than in the Teleosauria. The pro- 
portions of these bones, then, lead us to look for a stouter thigh and 
a shorter and broader foot than exist in the Cvrocodt/a. On the 
other hand, the single natural cast of a long and nearly straight 
bone, which I can only regard as an ungual phalanx, indicates a 
length of claw wholly foreign to the Crocodilian foot. 
The vertebra, whose more or less perfect natural casts have 
passed through my hands, all belong either to the dorsal, the sacral, 
or the caudal series. The impression of part of a sacral vertebra, to 
which I have already alluded, shows that in this region the structure 
of Stagonolepis was very similar to that of known Crocodz/ia; but the 
dorsal and caudal vertebra are remarkable, partly for the lateral 
constriction and inferior excavation of their centra, partly for the 
obliquity of the planes of their slightly concave anterior and 
posterior articular faces. These faces, in fact, are not perpendic- 
ular to the longitudinal axis of the centrum, but the anterior one 
looks a little downwards and forwards, the posterior, upwards and 
backwards. 
The neural arches were readily detached from their centra, and, 
where a separation has taken place, the previously co-adapted sur- 
faces of the centrum and the arch exhibit strong ridges and grooves, 
which mutually interlock. The edges of the posterior zygapo- 
physes meet inferiorly above the neural canal (which is deepest in the 
middle), forming a kind of inverted V. 
These and some other peculiarities of the vertebrae of Stagonolepis 
are all shared by the Teleosauri, and may be readily seen in many 
of the detached Teleosaurian vertebre in the British Museum; but 
the vertebre of the reptile under description present two remarkable 
