H. G. SEELEY ON A PURBECK ORNITHOSAURIAN. 465 



33. On cm Ornithosatjrian (Doratorhynchtts validus) from the 

 Purbeck Limestone of Langton near Swanage. By Harry 

 Govier Seeley, Esq., F.L.S., E.G.S., Professor of Physical Geo- 

 graphy in Bedford College, London. (Read May 26, 1875.) 



In the Christmas of 1868, when staying at Swanage, I was so for- 

 tunate as to obtain from a quarryman a portion of a large lower jaw, 

 in association with a long vertebra, which indicate an Ornithosaurian 

 animal of unusual size. Neck and jaw are parts to be expected in 

 close association ; but I see no reason to believe, or doubt, that the 

 bones pertained to the same individual, though in the absence of 

 conflicting evidence I shall be justified in attributing both specimens 

 to the same species. As with all my findings, these were deposited 

 in the Woodwardian Museum of the University of Cambridge, and 

 briefly noticed in 1869 in my ' Index to the Aves, Ornithosauria, 

 and Heptilia,' pp. 89, 90, the species being named Pterodactylus 

 macrurus. I now offer some further description of the remains, 

 and propose to place them in a new genus. The jaw cannot be 

 located in the genus Pterodactylus, because no evidence exists of 

 the occurrence in England of that genus (which, so far as I can 

 discover, has been found only in the Solenhofen slate, and is repre- 

 sented by animals of small size), and because no specimen of Ptero- 

 dactylus has the compressed, elongated, many-toothed, spear-shaped 

 jaw on which I found the genus Doratorhynchus. Pterodactylus 

 had the teeth in the jaw directed upward, and it is distinguished by 

 having also a tail as short as that of a rabbit or deer ; while this 

 specimen (if the vertebra is caudal, and if the tail may be inferred 

 from a single vertebra, five inches in length) would have had a tail 

 unusually long and of considerable strength, and it possessed a 

 flattened jaw, with teeth directed outward. The flattened jaw sug- 

 gests Cycnorhamphus suevicus of the Lithographic slate as an ally ; 

 but since the vertebra, as I shall presently show, may be cervical, 

 as may all those from the Cambridge Greensand which have been 

 regarded as caudal, no definite generic character can be drawn from 

 the vertebra alone. 



The lower jaw. 



The rami do not extend so far back as the articulation with the 

 quadrate bone. So much of the specimen as is preserved measures 

 12^ inches in length ; and, where fractured behind, the rami measure 

 2\ inches from side to side. The symphysis, beautifully preserved, 

 extends for 5 inches. The jaw, at its anterior termination, where 

 it expands a little, is nearly J inch wide, while at the posterior ter- 

 mination of the symphysis it is an inch wide. The anchylosed por- 

 tions of the rami are marked, as is usual, by a deep palatal groove. 

 The teeth were very small and close-set, and have all fallen from 

 their sockets. 



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