﻿THE 
  PHYLOGENY 
  OF 
  THE 
  ALUCITIDES. 
  

  

  121 
  

  

  resting 
  on 
  the 
  eosta 
  and 
  extending 
  down 
  to 
  a 
  point 
  just 
  within 
  the 
  end 
  of 
  the 
  fissure. 
  

   One 
  or 
  two 
  light 
  lines 
  cross 
  the 
  lobes 
  obliquely, 
  and 
  there 
  is 
  a 
  dark 
  spot 
  on 
  the 
  cell 
  

   a 
  little 
  before 
  the 
  middle 
  of 
  the 
  wing, 
  and 
  another 
  on 
  the 
  fold 
  still 
  nearer 
  the 
  base 
  

   of 
  the 
  wing. 
  The 
  hindwings 
  are 
  of 
  one 
  uniform 
  colour, 
  and 
  seldom 
  have 
  spots 
  or 
  

   lines 
  of 
  other 
  colours. 
  

  

  There 
  are 
  many 
  interesting 
  facts 
  relating 
  to 
  the 
  habits 
  of 
  the 
  

   Alucitids 
  which 
  might 
  be 
  noted 
  here. 
  In 
  their 
  hybernation 
  they 
  

   appear 
  to 
  be, 
  on 
  the 
  whole, 
  very 
  uniform 
  in 
  their 
  habits, 
  the 
  greater 
  

   number 
  hybernating 
  as 
  larvae, 
  and 
  falling 
  into 
  three 
  distinct 
  groups 
  : 
  

   (1) 
  The 
  Platyptiliicls 
  (sens, 
  lat.), 
  which 
  appear 
  mostly 
  to 
  hybernate 
  in 
  

   the 
  second 
  stadium. 
  (2) 
  The 
  Alucitids, 
  which 
  appear 
  mostly 
  to 
  hybernate 
  

   in 
  the 
  fourth 
  stadium. 
  (3) 
  The 
  exceptional 
  Leioptilids— 
  Adaina 
  (micro- 
  

   dactyla), 
  Hellinsia 
  (osteodactyla), 
  and 
  possibly 
  one 
  or 
  two 
  other 
  allies 
  — 
  

   which 
  hybernate 
  fully 
  grown. 
  The 
  first 
  group, 
  i.e., 
  those 
  that 
  hybernate 
  

   as 
  very 
  small 
  larvae, 
  are, 
  so 
  far 
  as 
  is 
  at 
  present 
  known, 
  all 
  in 
  their 
  

   early 
  larval 
  stages, 
  i.e., 
  preceding 
  hybernation, 
  internal 
  feeders, 
  boring 
  

   as 
  soon 
  as 
  hatched 
  into 
  the 
  foodplant 
  and 
  forming 
  a 
  hybernaculum 
  

   within 
  the 
  stem 
  when 
  fullfed 
  in 
  the 
  second 
  instar, 
  e.g., 
  Stenoptilia 
  ptero- 
  

   dactyla, 
  etc. 
  It 
  would 
  appear, 
  however, 
  that 
  the 
  Platyptiliines 
  do 
  not 
  so 
  

   fully 
  hybernate 
  as 
  some 
  of 
  their 
  congeners 
  — 
  Platyptilia 
  isodactylus 
  being- 
  

   inclined 
  to 
  feed 
  all 
  the 
  winter, 
  in 
  suitable 
  weather, 
  and 
  P. 
  gonodactyla 
  mak- 
  

   ing 
  a 
  start 
  in 
  the 
  very 
  earliest 
  days 
  of 
  spring, 
  etc. 
  — 
  yet 
  they 
  appear 
  to 
  go 
  

   into 
  winter-quarters 
  in 
  their 
  second 
  stadium. 
  The 
  second 
  group 
  has 
  

   larvae 
  that 
  are 
  external 
  feeders, 
  when 
  young, 
  and 
  most 
  of 
  these 
  appear 
  

   to 
  pass 
  the 
  winter 
  at 
  about 
  the 
  fourth 
  stadium 
  ; 
  the 
  young 
  larvae 
  appear 
  

   to 
  eat 
  furrows 
  into 
  the 
  leaves 
  without 
  making 
  an 
  actual 
  burrow, 
  and 
  

   hybernate 
  on 
  these, 
  in 
  some 
  instances, 
  e.g., 
  Porrittia 
  galactodactyla, 
  

   when 
  they 
  have 
  fallen 
  to 
  the 
  ground, 
  for, 
  strangely 
  enough, 
  they 
  

   mostly 
  feed 
  on 
  plants 
  that 
  die 
  down 
  in 
  the 
  winter. 
  The 
  third 
  group 
  

   is 
  exceptional, 
  Adaina 
  (niicrodactyla) 
  hybernating 
  as 
  a 
  fullfed 
  larva 
  in 
  

   the 
  cavity 
  that 
  will 
  become 
  its 
  puparium 
  in 
  spring 
  ; 
  Helliima 
  (osteo- 
  

   dactyla), 
  on 
  the 
  other 
  hand, 
  making 
  a 
  hybernaculum 
  among 
  the 
  foodplant, 
  

   but 
  apparently 
  leaving 
  this 
  (without, 
  however, 
  doing 
  any 
  more 
  feeding) 
  

   for 
  pupation 
  in 
  the 
  spring. 
  One 
  species, 
  Oidaematophorus 
  lithodactyla, 
  

   hybernates 
  in 
  the 
  egg-stage, 
  but 
  the 
  young 
  larva 
  is 
  developed 
  within 
  

   the 
  egg 
  very 
  shortly 
  after 
  the 
  latter 
  is 
  laid, 
  and 
  really 
  the 
  species 
  

   hybernates 
  as 
  a 
  young 
  larva 
  within 
  the 
  eggshell. 
  The 
  embryonic 
  

   larva 
  of 
  this 
  species 
  is 
  remarkable 
  in 
  having 
  a 
  highly 
  spiculate 
  skin- 
  

   surface 
  (resembling 
  those 
  of 
  other 
  species 
  in 
  their 
  second 
  or 
  third 
  

   instars), 
  not 
  that 
  no 
  other 
  newly-hatched 
  larva 
  is 
  spiculated, 
  tor 
  those 
  

   of 
  Adaina 
  niicrodactyla, 
  Herri 
  field 
  ia 
  tridactyla 
  (tetradactyla), 
  Adactylus 
  

   bennetii, 
  etc., 
  have 
  some 
  traces 
  thereof, 
  but 
  in 
  none 
  of 
  fchese 
  is 
  it 
  so 
  

   developed, 
  even 
  in 
  the 
  second 
  instar, 
  as 
  in 
  the 
  newly-hatched 
  larva 
  of 
  

   0. 
  lithodactyla. 
  There 
  appears 
  to 
  be 
  no 
  plume 
  that 
  hybernates 
  as 
  a 
  

   pupa, 
  but 
  two 
  allied 
  species 
  on 
  the 
  IMat\ 
  ptiliine 
  phylum 
  (Aiuh/>i/>tilia 
  

   cosmodactyla 
  and 
  A. 
  panctidactyla), 
  and 
  one 
  on 
  the 
  Alucitine 
  phylum 
  

   (Emmelina 
  monodactyla) 
  hybernate 
  as 
  imagines. 
  The 
  peculiar 
  res 
  ting- 
  

   habits 
  of 
  the 
  imagines, 
  e.g., 
  the 
  hanging 
  Platyptiliid, 
  the 
  doublefinger- 
  

   post 
  Emmelinid, 
  etc., 
  will 
  be 
  dea.lt 
  with 
  in 
  our 
  notes 
  on 
  the 
  imaginal 
  

   habits 
  of 
  the 
  various 
  species. 
  

  

  THE 
  PHYLOGENY 
  F 
  THE 
  ALUCITIDES, 
  

  

  The 
  position 
  of 
  the 
  Alucitids 
  and 
  then- 
  phylogenetic 
  relationships 
  

   are 
  problems 
  that 
  are 
  still 
  unsolved. 
  From 
  the 
  tune 
  of 
  lunne. 
  they 
  

  

  