﻿ETJCNEMIDOPHORINiE. 
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  and 
  without 
  any 
  ridge 
  connecting 
  each 
  with 
  the 
  next. 
  That 
  the 
  

   subdorsal 
  ridges 
  on 
  the 
  last 
  segments, 
  running 
  down 
  into 
  the 
  

   cremastral 
  pen, 
  are 
  a 
  reappearance 
  of 
  the 
  same 
  structure, 
  is 
  possible, 
  but 
  

   certainly 
  doubtful 
  ; 
  there 
  is 
  never 
  any 
  continuous 
  ridge 
  from 
  the 
  

   forward 
  to 
  the 
  posterior 
  portion. 
  In 
  the 
  Agdistid 
  pupae 
  some 
  traces 
  of 
  

   a 
  similar 
  structure 
  exist, 
  at 
  least 
  on 
  the 
  thorax, 
  and 
  are 
  marked 
  on 
  

   the 
  mesothorax 
  of 
  Herbertia 
  tamaricis, 
  for 
  example, 
  but 
  they 
  appear 
  to 
  

   be 
  more 
  correctly 
  regarded 
  as 
  the 
  pupal 
  representatives 
  of 
  the 
  larval 
  

   horns 
  and 
  processes, 
  as 
  connecting 
  ridges 
  are 
  wanting, 
  as 
  in 
  the 
  after- 
  

   warts 
  of 
  the 
  pupae 
  of 
  Amblyptilia. 
  When 
  we 
  turn 
  to 
  the 
  Alucitids, 
  we 
  

   find 
  structures 
  that 
  at 
  first 
  sight 
  appear 
  to 
  be 
  these 
  dorsal 
  ridges, 
  as, 
  

   for 
  example, 
  on 
  the 
  pupa 
  of 
  Ocendenia 
  septodactyla 
  (Menigianus), 
  where 
  

   the 
  fan 
  of 
  hairs 
  on 
  the 
  mesothorax, 
  especially 
  when 
  seen 
  from 
  above, 
  

   has 
  all 
  the 
  aspect 
  of 
  such 
  a 
  ridge 
  ; 
  when 
  looked 
  at 
  laterally, 
  it 
  is 
  seen, 
  

   however, 
  to 
  continue 
  so 
  far 
  as 
  hairs 
  arise, 
  i.e., 
  it 
  is 
  really 
  a 
  specially 
  

   formed 
  wart, 
  but 
  it 
  does 
  not 
  continue 
  beyond, 
  or 
  run 
  from, 
  segment 
  to 
  

   segment. 
  No 
  doubt 
  these 
  fan-shaped 
  warts, 
  the 
  after-warts 
  of 
  

   Amblyptilia, 
  etc., 
  are 
  all 
  expressions 
  of 
  a 
  readiness 
  of 
  the 
  skin 
  

   structures 
  to 
  assume 
  special 
  forms, 
  of 
  which 
  the 
  dorsal 
  ridges 
  are 
  one; 
  

   no 
  doubt 
  also, 
  in 
  a 
  sense, 
  they 
  are 
  similar 
  responses 
  of 
  this 
  readiness 
  

   to 
  similar 
  stimuli 
  ; 
  but 
  it 
  seems 
  highly 
  probable 
  that 
  they 
  are 
  separate 
  

   responses 
  in 
  separate 
  cases, 
  and 
  not 
  variations 
  of 
  one 
  original 
  ancestral 
  

   development 
  (Chapman). 
  

  

  So 
  far 
  as 
  the 
  three 
  constituent 
  groups 
  included 
  in 
  this 
  discussion 
  are 
  

   concerned, 
  their 
  general 
  characters 
  suggest 
  them 
  not 
  as 
  a 
  homogeneous 
  

   group, 
  like 
  the 
  Platyptiliines 
  or 
  Stenoptiliines, 
  but 
  as 
  separate 
  

   sections. 
  This 
  comes 
  out 
  particularly 
  in 
  the 
  genital 
  appendages. 
  This 
  

   could 
  be 
  met 
  by 
  making 
  each 
  the 
  centre 
  of 
  a 
  group 
  equal 
  in 
  value 
  to 
  

   the 
  Platyptiliinae 
  or 
  Stenoptiliinae, 
  as 
  we 
  have 
  done, 
  or 
  by 
  treating 
  

   them 
  as 
  three 
  separate 
  tribes 
  in 
  a 
  subfamily 
  Amblyptiliinae. 
  The 
  im- 
  

   portant 
  thing 
  is 
  to 
  get 
  the 
  differences 
  defined. 
  The 
  units, 
  at 
  any 
  

   rate, 
  have 
  no 
  close 
  alliance, 
  as 
  have 
  those 
  of 
  the 
  Platyptiliinae, 
  

   Stenoptiliinae 
  or 
  Oxyptilinae 
  ; 
  we 
  think 
  it 
  very 
  possible 
  that 
  a 
  

   sufficiently 
  detailed 
  knowledge 
  of 
  the 
  early 
  stages 
  of 
  a 
  greater 
  number 
  

   of 
  extra-British 
  forms 
  would 
  strongly 
  support 
  us 
  in 
  making 
  each 
  the 
  

   centre 
  of 
  a 
  subfamily 
  of 
  equal 
  classificatory 
  value 
  with 
  the 
  three 
  sub- 
  

   families 
  above 
  mentioned. 
  

  

  Subfamily 
  : 
  EucNEMiDOPHORiNiE. 
  

   Tribe 
  : 
  Eucnemidophoridi. 
  

   This 
  subfamily 
  is 
  much 
  more 
  distinctly 
  Platyptiliine, 
  than 
  are 
  the 
  

   two 
  succeeding 
  subfamilies. 
  So 
  much 
  is 
  this 
  so, 
  that 
  Bacot 
  considers 
  

   that 
  the 
  Eucnemidophoridi 
  would 
  be 
  perhaps 
  better 
  placed 
  as 
  a 
  tribe 
  of 
  

   the 
  Platyptiliinae 
  than 
  united 
  with 
  Amblyptiliinae, 
  but 
  the 
  differences 
  

   between 
  them 
  are 
  also 
  great, 
  and 
  so 
  we 
  separate 
  it 
  from 
  both. 
  In 
  the 
  

   larva 
  of 
  Eucnemidophorus, 
  the 
  depression 
  of 
  the 
  prothoracic 
  seutellum 
  

   is 
  hardly 
  noticeable 
  ; 
  the 
  primary 
  seta? 
  are 
  smooth 
  and 
  tapering 
  (some- 
  

   times 
  knobbed); 
  the 
  accessory 
  postspiraculars 
  are 
  marked 
  by 
  the 
  

   presence 
  of 
  slightly 
  better-developed 
  secondary 
  hairs, 
  the 
  skin-hairs 
  

   being 
  of 
  distinctly 
  Amblyptiliine 
  pattern. 
  In 
  this 
  stage, 
  Bacot 
  observes 
  

   that 
  there 
  is 
  a 
  strong 
  growth 
  of 
  secondary 
  skin-hairs, 
  and 
  that 
  the 
  

   Eucnemidophorid 
  Larva 
  presents 
  also 
  a 
  suggestion 
  that 
  the 
  accessory 
  

   postspiracular 
  tubercles 
  form, 
  as 
  it 
  were, 
  a, 
  sort 
  of 
  link 
  between 
  the 
  

  

  