47 



which is divided into two portions : a large and broad proximal portion , lying 

 in the sharp curve at the beginning of the third portion of the seminal vesicle; 

 and a small and narrower, but much longer distal portion. The ciliated canal 

 (Ci. C.I, recently discovered by Chun, opens into the distal end of the third 

 portion of the seminal vesicle. 



The appendix forms the small auricular fore lobe of the spermatophoric 

 gland. Its wall is plain , except that the epithelium of the small , widely-open 

 upper diverticulum forms a series of low longitudinal ridges. The distal end of 

 the appendix leads to the vas efferens , which extends., from a point slightly in 

 front of the distal end of the vas deferens , nearly to its proximal end. The 

 prostate or accessory gland is a large , flattened , broadly elliptical organ which 

 extends from the appendix nearly to the hind end of the gland , and lies inside 

 and below the veslcula seminalis. Its wall forms a series of narrow longitudinal 

 ridges which separate a series of deep , long and narrow pits , so that the gland 

 is of the simple tabular type. Its central lumen is quite large , and surrounds the 

 proximal part of its long tubular duct which opens into the appendix. 



The appendicular gland (Di) , a slender and relatively short blind tube 

 formed of columnar epithelium , is nearly one half as long as the entire gland 

 and extends backward inside the prostata from the distal end of the appendix. 



The ciliated canal (Plate II, Fig. 12, Ci.C.) is a small and long ciliated 

 tube whose wall, in prepared sections, is thrown into a few low longitudinal 

 folds. It begins near the hind end of the gland in a broad ciliated funnel whose 

 two divergent halves rest upon the adjacent surfaces of the accessory gland and 

 the third portion of the seminal vesicle , and it opens into or from the genital 

 sac , whose wall covers the gland. The canal extends forward between the acces- 

 sory gland and the third portion of the vesicula seminalis to the distal end of 

 the latter , which it enters obliquely. The function and morphological meaning 

 of the ciliated canal are alike doubtful. Marchand finds that its cilia beat away 

 from the sac and toward the gland . but its broad flaring funnel and its oblique 

 opening into the gland seem to indicate that they must carry fluid into the gland. 

 Marchand believes , however , that it is the distal portion of a second sexual 

 duct, whose proximal portion has been fused with the vas deferens, and that 

 it corresponds morphologically to the portion of the internal oviduct which is 

 buried in the wall of the oviducal gland , the latter being the homologue of the 

 genital pouch , whereas the nidamental gland corresponds to the vas efferens , 

 penis and spermatophoric gland , and has lost its primitive connection with the 

 proximally fused genital ducts. Bourne finds, in Sepia, that the ciliated canal 

 leads from the coelom to the spermatophoric gland, and this seems to favor 



