Abstract, 3 

 Resume, 4 



Acknowledgements, 5 

 Introduction, 7 

 History, 7 

 Materials, 12 

 Methods, 12 



Counting and measuring, 12 



Statistics, 12 

 Morphological results, 16 

 Biological data, 16 



Reproduction, 16 



Food, 19 



Size, age and growth, 19 



Habitat, 20 

 Transplantation experiments, 20 

 Discussion, 21 



Taxonomy, 21 



Nomenclature, 23 



Key, 24 



Distribution, 24 

 Origin of pygmy smelt, 26 

 Management implications, 27 

 References, 29 



