190 Prof. Frankland on the Origin of Muscular Power. 



below as A, B, C, and D, were employed in these experiments; 

 and each worked upon the wheel in alternate quarters of an hour, 

 resting in a sitting posture during the intervening quarters. 

 The period of actual daily labour was 3^ hours. The total ascent 

 per hour 2160 feet, or per day 1*432 mile. The following are 

 the results : — 







Treadweel Work. 



(E. Smith.) 







Weight in 

 kilogs. 



Ascent in 

 metres. 



Days 

 occupied 

 in ascent. 



External 

 work per- 

 formed in 

 metre- 

 kilogs. 



Total 

 nitrogen 

 evolved. 



Weight of 

 dry muscle 

 correspond- 

 ing to ni- 

 trogen. 



A 

 B 



C ' 

 D 



476 

 49 

 55 

 56 



23,045 

 23,045 

 20,741 

 20,741 



10 



10 



9 



9 



1,096,942 

 1,129,205 

 1,140,755 

 1,161,496 



grms. 

 171-3 

 174-5 

 168-0 

 1593 



grms. 



1101-2 



1121-7 



10801 



1024-3 



In these experiments the measured work was performed in the 

 short space of 3^ hours, whilst the nitrogen estimated was that 

 voided in the shape of urea in twenty-four hours. It will there- 

 fore be necessary to add to the measured work that calculated for 

 respiration and circulation for the whole period of twenty-four 

 hours. This amount of internal work was computed from the 

 estimates of Helmholtz and Fick as follows : — 



Internal Work. (Helmholtz and Fick.) 





Work 

 performed. 



Actual energy 

 required. 



Circulation of the blood during 24 T 

 hours at 75 pulsations per minute J 



Respiration for 24 hours at 12 pul- \ 

 sations per minute J 



metrekilogs.* 

 69,120 



10,886 

 Not determined. 



>> >> 



metrekilogs. 

 138,240 



21,772 

 Not determined. 



»» » 









80,006 



160,012 



Taking this estimate for internal work, the average results of 

 the treadwheel experiments may be thus expressed : — 



* Since making use of this number I find that Donders estimates the 

 work of the heart alone for twenty-four hours at 86,000 metrekilograms, a 

 figure which is higher than that used above for the combined work of cir- 

 culation and respiration. 



